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Consumption rate varies seasonally. Maximum rates usually occur in <br />spring or early summer (Johnson 1966a; Weithman and Anderson 1977; Diana <br />1979), following the spawning period. Weithman and Anderson (1977) observed <br />peak feeding at temperatures between 15 and 18° C for yearling northern pike <br />held under experimental conditions; there was a summer reduction in consump- <br />tion rate between May and September highs. <br />Reproduction <br />Northern pike spawn in spring, shortly after ice-out, when the water has <br />warmed to 8 to 12° C (Embody 1918; McNamara 1936; Clark 1950; Fabricius and <br />Gustafson 1958; Franklin and Smith 1963; June 1971; Priegel and Krohn 1975). <br />Movement to spawning grounds often begins before all ice has melted. Upstream <br />spawning migrations out of lakes in Michigan (Carbine 1942) and Minnesota <br />(Franklin and Smith 1963) occurred as soon as there was sufficient clearance <br />between inshore ice-and the bottom to permit passage. Robertson (1969) <br />observed that upstream movement of northern pike was interrupted when the <br />water temperature dropped below 5.5° C. Isolated spawning may occur at tem- <br />peratures less than 8° C (McNamara 1936; Clark .1950; Schryer et al. 1971). <br />Most females have completed spawning by the time temperatures exceed 13° C <br />(Schryer et al. 1971). <br />Spawning occurs over vegetation in areas of calm, shallow water <br />(Williamson 1942; Clark 1950; Fabricius 1950). There is a strong tendency to <br />migrate up tributaries to flooded marshes, wetlands, or shallow pools among <br />both lake (Carbine and Applegate 1948; Schultz 1955; Robertson 1969; Koshinsky <br />1979) and river (Harrison and Hadley 1978) populations. Flooded terrestrial <br />vegetation (McCarraher and Thomas 1972) and shallow, weedy bays or backwaters <br />may also be used (Jarvenpa 1962a; Frost and Kipling 1967). The absence of <br />inundated vegetation can inhibit or delay spawning (Fabricius 1950; Fabricius <br />and Gustafson 1958). The timing of spawning has been linked to water level <br />changes (Fabricius 1950; June 1970, 1971, 1977). <br />Spawning groups, consisting of a female plus one, or more typically, <br />several, males move continually. Only a few eggs, 5 to 60 according to <br />Svardson (1949), are released at any one spot (Svardso~ 1949; Clark 1950; <br />Fabricius and Gustafson 1958). Gametes are broadcast (Fabricius and Gustafson <br />1958), and no parental care is provided (Eddy and Underhill 1974). Eggs <br />adhere to vegetation (Fabricius and Gustafson 1958; Frost ,and Kipling 1967). <br />Both .laboratory (Fabricius and Gustafson 1958) and field (Koshinsky 1979) <br />observations indicate that spawning occurs in distinct series of mating acts, <br />between which the fish are stationary, presumably to rest.- Spawning groups <br />may cover considerable distances before all eggs are released (Koshinsky <br />1979). Distances between egg releases are likely to be greater when the <br />distribution of spawning habitat is patchy (Fabricius and Gustafson 1958). <br />Northern pike do not spawn at night (Clark 1950; Fabricius and Gustafson <br />1958). Spawning may be interrupted by cold weather, water ]evel drawdowns <br />(Clark 1950; Fabricius and Gustafson 1958; June 1970), strong wind, or rain <br />(Koshinsky 1979). Prolonged interruptions may result in resorption of eggs <br />(June 1970). <br />~~ <br />Q <br />3 <br />