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provinces and were grouped in more than one investi- <br />gational category and in more than one land use. <br />Algal Communities <br />Ten investigations between 1962 and 1987 <br />incorporated algal-community information <br />(table 1, fig. 2). Sites in these studies were located <br />in all predominant land-use categories and in both <br />physiographic provinces (table 1, fig. 1). The algal <br />data in these investigations were presented mostly as <br />taxonomic identifications for the stream segments. <br />The summaries of these investigations indicate that <br />algal data are limited throughout the basin, but that <br />algae are very useful indicators of water chemistry. <br />Land use can affect types, numbers, and <br />diversity of algae. Mining areas contain different taxa <br />than that present in agricultural areas. For instance, <br />in the UCOL study unit, investigations done in mining <br />areas commonly contain certain species of green <br />algae and diatoms because of their tolerance to acidic <br />conditions and metals. Agricultural areas commonly <br />contain blue-green algae as the predominant species <br />because of its tolerance to higher concentrations <br />of nitrogen from fertilizers (Stephen Porter, <br />U.S. Geological Survey, oral common., 1996). <br />Macroinvertebrate Communities <br />Fifty investigations between 1938 and 1995 <br />included macroinvertebrate-community information <br />(table 1, fig. 3). Sites in these studies were located in a <br />variety of land uses and in both physiographic <br />provinces (table 1, fig. 1). These investigations <br />presented the data as taxonomic identifications or as <br />numbers of organisms. Some of the investigations <br />presented both types of data. The summaries of these <br />investigations indicate there is an abundance of <br />information on macroinvertebrate communities <br />throughout the UCOL study unit. <br />Land use can have direct effects on the <br />composition of macroinvertebrate communities. The <br />use of indicator species for water-quality analysis is <br />very common in studies of macroinvertebrate <br />communities, especially when determining the water <br />quality from mining effects. Macroinvertebrate- <br />community composition can be different in mining <br />areas than in agricultural areas. Mining areas <br />commonly are depleted of aquatic organisms, but <br />can contain metal-tolerant species. Agricultural areas <br />commonly contain species, such as aquatic worms <br />and leeches, that are tolerant oforganic-carbon enrich- <br />ment, sedimentation, and low dissolved-oxygen <br />concentrations (Stephen Porter, U.S. Geological <br />Survey, oral common., 1996). <br />Fish Communities <br />Eighty-nine investigations summarized fish <br />community or population information between 1938 <br />and 1995 (table 1, fig. 4). Sites in these studies were <br />located in a variety of land uses and in both physio- <br />graphic provinces (table 1, fig. 1). The information on <br />fish was presented as taxonomic identifications or as <br />numbers offish. Some of the investigations presented <br />both types of data. The summaries of these investiga- <br />tions indicate that information on fish communities is <br />extensive throughout the UCOL study unit. <br />Fish communities are an important issue in the <br />UCOL study unit; the study unit contains four fishes <br />presently listed as endangered by the U.S. Fish and <br />Wildlife Service (USFWS) (Behnke and Benson, <br />1980): the Colorado squawfish (Ptychocheilus <br />Lucius), the humpback chub (Gila cypha), the bonytail <br />chub (Gila elegans), and the razorback sucker <br />(Xyrauchen texanus). In addition, recreational fishing <br />in the study unit is important to many anglers. <br />Historically, native fauna consisted of the <br />following minnows and suckers in addition to those <br />fishes listed above: roundtail chub (Gila robusta), <br />flannelmouth sucker (Catostomus latipinnis), speckled <br />dace (Rhinichthys osculus), kendall warm springs dace <br />(Rhinichthys osculus thermalis), and bluehead sucker <br />(Catostomus discobolus} (Tyus and others, 1982). The <br />current fauna of the Colorado River is dominated by <br />exotic species. All of the big river fish that were <br />native to the area, except for the speckled dace, <br />flannelmouth sucker, bluehead sucker, and roundtail <br />chub, face extinction. The cause of the disappearance <br />of some of the native species is due to loss of habitat <br />and competition from other species (Ward and others, <br />1986). Native fish in the upper reaches of the basin <br />included mountain sucker (Catostomus platyrhyn- <br />chus), mountain whitefish (Prosopium williamsoni), <br />mottled sculpin (Cottus bairdi), and Colorado <br />cutthroat trout (Salmo clarki pleuriticus). <br />20 Summary of Biological and Contaminant Investigations Related to Stream Water Quality and Environmental Setting in the <br />Upper Colorado River Basin, 1938-95 <br />