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1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />1 <br />The basic model structure is: <br />Nom, =Nr+R-M+1-E <br />for each river section where: Nt is the number of individuals of a given age at a <br />specific time, R is the recruitment from the previous age class, M is the number of <br />individuals that die for that age class, 1 is immigration from the upstream section for the <br />age class, and E is emigration out of the section for that age class above carrying <br />capacity. <br />1. The sequence begins with spawning adults (initial population) occurring at <br />designated spawning areas. The number of spawning adults is equal to the sum of all <br />adults for all sections above and below the spawning area within the subpopulation <br />boundaries (e.g., Middle Green, Lower Green, and Colorado River for Colorado <br />squawfish). After spawning, the adults move back to the reach from which they <br />originated, to the adult habitat reaches identified and assess the number of fish already <br />present relative to the carrying capacity for adults. If the number of adults already <br />present equals carrying capacity, then the adults continue to the next available adult <br />habitat section. If the carrying capacity is not exceeded, then all adults remains in their <br />original habitat section. For each habitat segment that the adults move through from <br />the spawning bed, a proportion can die based upon an adult mortality rate for dispersal. <br />The model is set for zero mortality due to movement at this time. <br />2. The adults in the spawning beds produce a number of eggs per female (set by a <br />10 <br />