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The basic model structure is: <br />N~,=Nt+R-M+1-E <br />for each river section where N=number of individuals of a given age, R is the <br />recruitment from the previous age class, M is the number of individuals that die for that <br />age class, / is immigration from the upstream section for the age class, and E is <br />emigration out of the section for that age class above carrying capacity. <br />1. The sequence begins with spawning adults (initial population) occurring at <br />designated spawning areas. The number of spawning adults is equal to the sum of all <br />adults for a!I sections below the spawning area within the subpopulation boundaries <br />(e.g., Middle Green, Lower Green and Colorado River for Colorado squawiish). After <br />spawning, the adults move downstream to the adult habitat reaches identified and <br />assess the number of fish already present relative to the carrying capacity for adults. If <br />the number of adults already present equals carrying capacity, then the adults continue <br />to the next adult habitat section. If the carrying capacity is not exceeded, then a <br />number remains so that the carrying capacity is equaled and the remaining adults <br />continue downstream. For each habitat segment that the adults have to move through <br />in moving downstream from the spawning bed, a proportion die based upon an adult <br />mortality rate for dispersal. This means that the further an adult has to move <br />downstream, the less likely it is to survive. <br />2. The adults in the spawning beds produce a number of eggs per female (set by a <br />9 <br />