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MODEL DEVELOPMENT <br />9riginal Model Development <br />We have generated a generalized, compartmentalized life-history model for Colorado <br />River Fishes. The original model (Appendix I) was written in using the Mathcad <br />software package as a stochastic population model based on the logistic growth model <br />(refs here). The model is spatially explicit to allow the exploration of subpopulation <br />dynamics using geomorphic units as the spatial units (Table 1). The model is also <br />age-specific to identify potentially limiting life stages. Important input variables for <br />geomorphically distinct river sections (e.g., Yampa R,, Green R., Colorado R.) include: <br />the number of eggs and larvae, young-of--the-year, age 1, age 2, age 3 and adult (>4 <br />yrs old) fish per river section. Important parameters included are: reproductive effort, <br />both in terms of numbers of young and where spawning occurs, downstream drift <br />dynamics of YOY fish determined by carrying capacity, mortality estimates with inter- <br />annual variation for each age class (listed above); and estimates of carrying capacities <br />for each age class within each river segment, and sex ratios of mature adults. <br />4 <br />