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<br />preceding fall. Larger and more age-O fish in fall resulted in larger and more age-l fish in spring, <br />which suggests that overwinter size-selective pressures were relatively consistent from year to <br />year. The fact that reduction in CPE from fall to spring varied from year to year indicates the <br />magnitude of variation in year class strength overrode any effect of overwinter mortality. Two <br />important conclusions can be drawn from this finding: I) year class size and length in fall may <br />override effects of overwinter size-selective mechanisms in dictating year class size and length in <br />spring and 2) current sampling in fall and spring are representative of actual conditions. <br />Our study further suggests that Flaming Gorge Dam has not altered the length of the <br />growing season as measured by DD based on growth potential indices. A similar finding was <br />reported by Kaeding and Osmundson (1988). They found that impoundment had not altered the <br />temperature regime of the middle reaches of the Colorado River and suggest that dams in the <br />lower Colorado River basin prevent Colorado pikeminnow from using warmer downstream <br />reaches (below Glen Canyon Dam). Yet, if this theory were true, one would expect age-O <br />Colorado pikeminnow to be larger in lower reaches of the Green River than in middle reaches <br />rather than being the same as noted in this and other studies (Tyus and Haines 1991). <br />It was unexpected that year class size and mean length in fall was not related to DD, <br />summer flows or timing and magnitude of peak discharge in the preceding spring. Possible <br />explanations for this may be that factors other than temperature and flow dictate number and size <br />of fish in fall and their respective condition overwinter. Survival and growth may be limited by <br />biotic factors such as predation by and competition with native and nonnative fish species before <br />winter. This hypothesis concurs with Bestgen (1997) who found biotic factors to be more <br />important than abiotic factors in determining fall recruitment of age-O Colorado pikeminnow, <br />which seems more likely than lower growth rates in the lower Green River. The lack of a size <br />difference between lower and middle age-O and age-l Colorado pikeminnow (despite a <br />substantial difference in temperature) could be the result of differences in habitat quantity and <br />quality between the two areas or may simply be an artifact of better survival of fish spawned later <br />in the season (when red shiner numbers decline) in the lower Green River (Bestgen 1997). Why <br />a similar relationship would not occur in the middle Green River is unclear. A loose association <br />of year class length and DD in the lower Green River and not in the middle Green River suggests <br />that limiting factors in the two reaches may be different. More information on ecology of and <br />predation by red shiners and other nonnatives is necessary to elicit this puzzling relationship. <br /> <br />CONCLUSIONS <br /> <br />Overall, scale analysis proved useful in determining the relationship between length and <br />overwinter mortality and growth; however, our age-O scale data was limited to several sample <br />sites within one year. Such data limitations precluded extensive statistical analysis and cautioned <br />that any conclusions be pending further investigations and validation of our findings. To validate <br />this relationship, we recommend that scale collection become a standard procedure for Inter- <br />agency Standardized Monitoring and that data for future analysis include a greater temporal and <br />spatial scale. In particular, this analysis should be evaluated for data representing more than one <br />year-class of age-O fish because of the inherent variability of year-class strength and size <br /> <br />13 <br />