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none were recaptured among years (two individuals within the same year). Non-existent captures <br />among years would cause the survival rate function to decline to near zero for the large fish <br />rather than increase as it did for the 1991 to 1999 data. Non-existent recaptures of large <br />individuals suggested that they were either very difficult to capture because they learned to avoid <br />the capture gear, used habitat where capture gear was not effective, or that some individuals died. <br />Lower survival rates of Colorado pikeminnow in the Green River Basin in the period 2000 <br />to 2003, particularly for larger-bodied fish, may be due to several factors including negative <br />effects of handling (including tagging) or electrofishing, stress-induced mortality due to low <br />streamflow levels in drought years 2002 and 2003, expanding populations of introduced <br />predaceous fishes such as northern pike, or a combination of these and other yet undiscovered <br />factors. Although population declines appeared consistent with increased mortality due to effects <br />of electrofishing or handling, we did not find support for that hypothesis. The number of <br />Colorado pikeminnow mortalities that we observed as a direct result of capture during this study <br />were small (N = 6 over the entire study), so high levels of electrofishing-induced or handling- <br />induced mortality were not evident. Colorado pikeminnow recaptured in multiple passes within a <br />year generally appeared in good condition, with no externals signs of damage (e.g., bruising), <br />handling-induced disease (e.g., fungus), or other ill effects. We have no data to address effects of <br />delayed mortality on Colorado pikeminnow, except that electrofishing-captured fish implanted <br />with radio transmitters for use in telemetry studies generally have high survival (Tyus 1990; <br />Kitcheyan and Montagne 2005). <br />In support of observations of healthy fish captured and recaptured in the field, independent <br />evaluations did not suggest electrofishing was harmful to Colorado pikeminnow. Controlled <br />laboratory studies (Meismer 1999) found minimal soft-tissue and vertebral damage from the <br />48 <br />r <br />r