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of recapture occasions (e.g., minimum of 5) are needed to detect differences in capture <br />probabilities among groups of animals in the same population. We also explain much individual <br />heterogeneity by including the covariate TL in other analyses. <br />Survival and finite population rate of change models.-Jolly-Seber type models <br />(recaptures only and Pradel's survival and population rate of change models) in program MARK <br />(Cormack 1964; Jolly 1965; Seber 1965; Pradel 1996; White and Burnham 1999) were used to <br />estimate apparent survival (1991 to 1999) and population rates of change (x,,1991 to 2003), <br />respectively, for Colorado pikeminnow captured in the Green River Basin. Apparent survival <br />rates (S) are the joint probability of a fish surviving from one year to the next and remaining in <br />the population available for capture. In other words, estimates from these models do not <br />distinguish a fish that died in the study area from one that survived and moved from the study <br />reach to an unsampled reach. Because of that, survival rates obtained from ISMP data are likely <br />conservative (low) because capture data were gathered from reaches that represent only about <br />23% of the basin. Thus, Colorado pikeminnow that move out of reaches sampled by ISMP and <br />are never recaptured would be treated as mortalities. These models are also not able to <br />distinguish if fish that were previously captured were able to avoid subsequent recapture by some <br />behavioral change mechanism. Such a behavioral change would result in reduced capture <br />probability and lower apparent survival rates. Assuming that no behavioral change occurred, <br />survival rates obtained from 2000-2003 during abundance estimation sampling likely approached <br />true survival, because nearly all the Green River Basin habitat where Colorado pikeminnow <br />likely occurred was sampled. The goal of survival analyses was to determine if a composite, and <br />perhaps conservative, survival rate from 1991 to 1999 was different than survival in the 2000 to <br />23