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outcome of these environmental and biotic changes for the highly endemic fish fauna of <br />Colorado River Basin has been dramatic: two of the 35 native species in the basin are extinct, an <br />additional 18 are federally listed as threatened or endangered or are very rare, and most others are <br />listed by one or more basin states as declining (Stanford and Ward 1986; Carlson and Muth <br />1989; Bezzerides and Bestgen 2002; Mueller and Marsh 2002). Colorado pikeminnow is <br />federally listed as endangered under the Endangered Species Act of 1973, as amended (U. S. Fish <br />and Wildlife Service 1974). <br />Colorado pikeminnow, endangered bonytail Gila elegans, humpback chub Gila cypha, <br />and razorback sucker Xyrauchen texanus are the focus of the Recovery Implementation Program <br />(RIP) in the Upper Colorado River Basin (Wydoski and Hamill 1991). Recovery goals for <br />Colorado pikeminnow provide criteria that must be achieved before downlisting or delisting can <br />be considered (U.S. Fish and Wildlife Service 2002). The recovery criteria include demographic <br />attributes that describe the required number of sub-basin populations and individuals (adults and <br />juveniles) within each population, and recovery factor criteria that are directly linked to <br />management actions and tasks needed to minimize or remove threats. <br />Maintenance of a metapopulation is central to Colorado pikeminnow recovery, and <br />demographic criteria require "a genetically and demographically viable, self-sustaining <br />population in the Green River sub-basin; and self-sustaining populations that meet or exceed <br />estimated carrying capacity either in only the upper Colorado River sub-basin, or in both the <br />upper Colorado River sub-basin and San Juan River sub-basin". For consideration of <br />downlisting or delisting, specific demographic criteria for the Green River sub-basin portion of <br />the Colorado pikeminnow metapopulation require that, over the specified monitoring period, (1) <br />trends in abundance estimates for adults (Z 450-mm total length, TL) in both the middle and <br />10