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similar to that of the squawfish in the main river <br />channels of the Colorado and Green rivers. Chubs <br />were restricted to swift, deepwater areas, mainly in <br />canyons, and did not occur far up tributary streams. <br />The greatest known concentration of this species now <br />occurs in the lower few miles of the Little Colorado <br />River, in the Grand Canyon area of Arizona. Perhaps <br />the releases of cold water from Lake Powell have <br />forced most of the humpback chubs from the main Colo- <br />rado River in Grand Canyon into the warmer Little <br />Colorado. <br />In the upper basin, humpback chubs occur sporadi- <br />cally in the Colorado River up to Palisade, Colo- <br />rado, about 10 miles above Grand Junction. The <br />greatest concentration occurs in the Black Rocks area <br />of Ruby Canyon, about 25 miles below Grand Junction, <br />where turbulent flows create a pool almost 60 feet <br />deep at low water levels. In the Green River, the <br />humpback chub occurs below the mouth of the Yampa, <br />and is concentrated in the Desolation Canyon area. <br />It has been recorded from the lower Yampa River in <br />Dinosaur National Monument. <br />Causes of Decline <br />Because the humpback chub had a restricted dis- <br />tribution and thus was always relatively rare, there <br />is not much evidence of a decline except where reser- <br />voirs were constructed. The deepwater habitat <br />favored by this species is not easily sampled by <br />standard methods of fish collecting. As sampling <br />techniques improve and more is learned about humpback <br />chubs, more populations probably will be discovered. <br />The most abundant known population, in the Little <br />Colorado River, was not discovered until 1975. <br />There has been considerable concern that the <br />humpback chub may lose its identity as a result of <br />hybridization with bonytail and roundtail chubs. It <br />now seems probable that most of the specimens formerly <br />believed to be hybrids actually showed only normal <br />variation in the degree of hump development. Some <br />specimens, however, probably are hybrids. The bony- <br />tail chub now is so rare that it can be discounted as <br />a significant source of possible hybridization. The <br />roundtail chub, however, is common in the Colorado <br />River in Colorado and occurs with the humpback chub <br />in Ruby Canyon, where some intermediate (hybrid?) <br />specimens have been taken. The roundtail chub is <br />absent or occurs rarely in humpback chub habitat in <br />the Green River or in the Little Colorado River. Thus, <br />overall, the threat to the integrity of the humpback <br />chub species from hybridization is probably not as <br />great as was once believed. <br />The deepwater areas preferred by humpback chubs <br />are also a preferred habitat for non-native channel <br />catfish. Large populations of catfish and carp share <br />the Ruby Canyon habitat with the humpback chub. <br />Prospects for the Future <br />A Recovery Plan has been written for the hump- <br />back chub, but, as with the squawfish, the main <br />emphasis was placed on hatchery propagation as the <br />only clearly defined technique to increase abundance. <br />Humpback chubs have been transported to the Willow <br />Beach National Fish Hatchery, Arizona, for an attempt <br />at artificial propagation. It is likely that addi- <br />tional populations will be found when more of the <br />deepwater canyon areas in the upper basin are more <br />thoroughly sampled. Fishermen can be of assistance <br />in this regard by reporting catches of humpback chubs. <br />Good humpback chub habitat is also good channel cat- <br />fish habitat, and the chub can be caught on the same <br />bait often used for catfish. Humpback chubs must, <br />of course, be released, but the Colorado Division of <br />Wildlife should be notified of the catch, particu- <br />larly if it is outside of the Ruby Canyon area of <br />the Colorado River. A documented angler's catch <br />(with a photo, if possible) may provide new distri- <br />bution records and lead to the discovery of new <br />populations of this rare fish. <br />A humpback chub preservation and restoration <br />program is yet to be started, but it will probably <br />consist of the identification of all areas where <br />populations still occur, so that the present environ- <br />mental conditions in those areas can be maintained. <br />It would be extremely difficult to establish humpback <br />chubs where they do not now exist. They may now <br />inhabit all suitable areas where self-sustaining <br />populations can be maintained under present environ- <br />mental conditions. The outlook is not encouraging <br />for expanding the distribution and abundance of <br />humpback chubs by establishing new self-sustaining <br />populations. Their habitat requirements are highly <br />restrictive. Possibilities should be looked for, <br />however, where deep channel areas have been created <br />by bridge or highway construction, forming suitable <br />habitat beyond the present limits of distribution. <br />In such situations, introduction of the humpback chub <br />might result in the successful establishment of a new <br />population. Valuable information could be obtained <br />from experimentation designed to establish new popu- <br />lations. There is little doubt that the humpback <br />17