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perforated treatment, chlorophyll a levels did not detect trophic cascade effects at the <br />phytoplankton level as is so often in the case in lentic systems (Carpenter et at. 1987, Johannes <br />et at. 1989, McQueen 1990, Shapiro and Wright 1984, but see McQueen et at. 1989). It is <br />somewhat surprising that there was no detectable effect of invertebrate predators on <br />oligochaetes since both predatory and omnivorous chironomids (such as TanXpus and <br />Chironomus), as well as corixids of the genus Tricorixa, are known predators of Oligochaetes. <br />Loden (1974) noted that oligochaetes often break into two pieces when in the grasp of a <br />predator. Given the regenerative abilities of oligochaetes, this would suggest that oligochaete <br />biomass may decrease in the presence of predators while their overall density remains about <br />constant (Wisniewski 1978). <br />iversi <br />As shown in Table 5, chironomids appear to be more diverse in control and perforated <br />areas. These results, coupled with the fact that all of the uncommon genera found were taken <br />from the control and perforated treatments, suggest that fish predation helps maintain the <br />diversity of chironomids in backwaters. This is consistent with other studies that have shown <br />that predation lessens the impact of competition and even prevents the local extinction of some <br />benthic invertebrates (Connell 1961, Paine 1966, Menge and Sutherland 1976). <br />Implications <br />The Colorado squawfish, North America's largest minnow (cyprinidae), now occupies <br />less than 25% of its historical range and is listed as endangered by the U.S. Fish and Wildlife <br />Service (Karp and Tyus 1990). While several factors have been implicated in the decline of <br />the Colorado squawfish, the rarity of juveniles (70-400 mm total length) suggests that the <br />11