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Sew <br />~. <br />research facilities for laboratory )studies. Fish stocking programs can benefit <br />management for recovery, but only in coordination with a comprehensive program <br />in which life requirements of target species are well understood, and these <br />requirements provided. If habitats have not improved, reintroduction of fishes <br />`into them should be considered only stop-gap measure (Rinne et al. 1986). i ~.`-Z~~~~~~'~ <br />p~diec,~~ev.. ~-~4~~. iw Y.ql~~~ ~ ~ w~,~. G~Q,~n4,~'~c. r~~o~kt VM~L~' ~kj~ :~l"Cw ~r~w.2~r caN,s~~2Ti~ lf~ <br />Stocking of Colorado squawfish for recovery has not been attem~ed for qNy <br />the Green River basin because of the naturally reproducing population there. 5~-0 ~~iµS <br />Re-introduction of Colorado squawfish into areas of the lower Colorado River ~-,~,~~-~l, <br />basin from which they have been extirpated, was initiated in 1985 by the <br />Arizona Game and Fish Department, and USFWS- Dexter National Fish Hatchery <br />(Hendrickson and Brooks 1987). It is too soon to determine-the success of that <br />endeavor, but it is doubtful if stocking alone will be sufficient to recover <br />the squawfish there. Indeed, the complex life cycle of the Colorado <br />squawfish, in which there are spatial separations of life stages, ~ ~- <br />incorporation of an energetically-costly migratory behavior, and other ~ov ~^- <br />components (Tyus 1986, 1990) suggests that stocking programs can only be ~ ~~ ~,~ <br />successfulif life history attributes are understood and considered. ~ ~ ~~~5 <br />Stocking of razorback suckers have been initiated in the Green River, ~a~ <br />but as yet this is only being done to augment a non-recruiting population ~~~ \~ v--- <br />(Lanigan and Tyus 1988, Severson et al. 1989). ~1 <br />Non-native Species and Sportfishing ~~v~~l~a~w~- <br />Stocking of nonnative, warmwater species ha bee reduced in the upper <br />Colorado River basin recent years, and under the R would not occur in <br />watersheds occupied by listed fishes. The reverse of stocking, elimination or <br />removal of non-natives, is worth considering, but this issue has not been <br />meaningfully addressed. <br />Introductions of non-native fishes into habitats occupied by native <br />fishes is not usually accompanied by a positive affect on native fishes, <br />especially if habitats have been altered. Direct effects include: Elimination, <br />reduced growth and survival, changes in community structure, and no effect <br />(Moyle et al'. 1986). In addition, fish are usually introduced into habitats <br />that are changing due to the effects of man, making it difficult to determine <br />if habitat change, competition, or both, are responsible for declines that are <br />noted in the native fauna. This already complex issue is further confounded by <br />past introductions into waters in which species interactions were not well <br />understood, and were usually not subsequently studied. Thus, impacts of <br />introduced fishes on native populations are seldom understood. Because it is <br />difficult to assess such impacts, and the effects are usually inferred from <br />major alterations of species composition. Competition between native and <br />introduced forms is difficult to observe experimentally, and perhaps <br />impossible to prove in natural riverine habitats. <br />The introduction of non-native fishes in the Colorado River has been <br />documented by many, and some data on their present status provided by Minckley <br />(1982) and Tyus et al. (1982). Competition of these non-natives with Colorado <br />squawfish has been proposed by various workers (Behnke and Bensen 1983; Holden <br />and Wick 1982; Osmundson 1987). Predation of introduced fishes on razorback <br />7 <br />