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<br />long as the physical habitat is appropriate for maintenance of the native prey <br />population, it is probably adequate for adult Colorado pikeminnow. <br />~ It is of considerable significance to the recovery effort that the preferred prey items are <br />less abundant in the UCR below Palisade. Adult Colorado pikeminnow may be <br />congregating in the 15-Mile Reach simply because it receives input of prey species <br />from upstream reaches and because it is as close as those fish can get to habitat that <br />contains the preferred prey. The possibility that adult Colorado pikeminnow abundance <br />is prey-limited in the 15-Mile Reach has significant ramifications for the recovery effort. <br />~ It suggests that, unless food supply can be increased, improvements to physical habitat <br />may not do much to increase the number of fish present. The area with suitable prey <br />for the large adult pikeminnow will be extended greatly when barriers to Debeque <br />Canyon are made passable. <br />~ Adult migration and spawning habitat. Lack of access to spawning grounds has been <br />implicated in the decline of the Colorado pikeminnow (Joseph et al. 1977, Tyus 1984). <br />Little is known, however, about the historical distribution and abundance of spawning <br />sites. Spawning habitat has been located in the Yampa, Green, and San Juan rivers, <br />and the presence of larvae shows that spawning also occurs in the UCR and the lower <br />~ Gunnison River. Habitats in the Yampa and San Juan rivers appear to conform to a <br />specific geomorphologic profile. The hydraulic and sedimentologic conditions <br />necessary for the creation and maintenance of habitat with those characteristics have <br />been defined recently by studying bar-forming events at two Colorado pikeminnow <br />spawning sites in the lower Yampa River (Harvey and Mussetter 1996, Harvey et al. <br />~ 1993). At present, there is no obvious reason to suspect that the quality or quantity of <br />spawning habitat in these three rivers is limiting reproduction of the pikeminnow. <br />Less is known about spawning habitat in the UCR, in part because there have been <br />only limited observations of adult pikeminnow occupying spawning ground, and no <br />running ripe females have been collected (D. Osmundson, personal communication. <br />~ Pikeminnow continue to spawn in the UCR despite the fact that some of the possible <br />spawning sites have been severely altered by land and water development (Valdez et <br />al. 1982, McAda and Kaeding 1991 b). To the extent that habitat in the UCR may <br />conform to the geomorphic profile that has been established for other spawning sites, it <br />may be possible to infer the location of the habitat. Supplementary data on the <br />~ distribution of larvae and ripe adults may help narrow the possibilities for spawning <br />sites. It will be very difficult to determine if spawning habitat may limit reproduction or <br />recruitment in the UCR until the location of the habitat is confirmed. In addition to <br />possible spawning sites in the Grand Valley and in the lower Gunnison River, there is <br />physical habitat in Debeque Canyon that appears to match the proposed <br />~ geomorphologic profile (see Anderson 1996). However, the site in Debeque Canyon is <br />not yet accessible to adults in the UCR because there are barriers to migration. <br />Adult population size. The minimum size required for maintaining a "viable° natural <br />population of any of the endangered fishes should be an important consideration for <br /> <br />30 <br /> <br />