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<br />Despite legitimate concerns about the extent to which present-day field studies of rare <br />species will provide an accurate representation of their life history requirements, there is <br />no alternative but to make the best use of such information. Field studies conducted <br />where a species is relatively abundant, and where the habitat is altered least (all habitat <br />~ now occupied by the endangered fishes has been altered some), are most likely to <br />provide an accurate view of life history requirements (discussed by Tyus 1992). The <br />optimal remaining habitat is closest to the conditions in which the native species <br />evolved, and presumably are the conditions in which the species are most likely to <br />maintain adaptive advantage over introduced species. Field studies in the UCRB have <br />~ tended to be very localized in focus, with the result that populations (or subpopulations) <br />of one species are treated almost as if they were separate species. A narrow <br />geographic focus for field studies can lead to a fragmented view of recovery needs. <br />Although local adaptations can and do occur, recovery efforts in general could benefit <br />from generalizations about and synthesis of life history needs, leading to a conceptual <br />~ model of life history. <br />The Green River system in the upper basin has long been identified as perhaps the <br />most suitable location to determine management measures necessary for recovery of <br />Colorado pikeminnow and razorback sucker, because it supports the largest riverine <br />~ populations of these species and because habitat in a major tributary, the Yampa River, <br />remains largely unaltered. Flow in the mainstem of the Yampa River is regulated little <br />and access to upstream habitat has not been blocked. The Grand Canyon in the <br />vicinity of the Little Colorado and Colorado rivers is the logical place to study the life <br />history needs for the humpback chub, because there is a large population sustained by <br />. natural recruitment. Information obtained from the more "natural° locations can be <br />supplemented, albeit cautiously, with observation from sites where habitat has been <br />altered, but where the species is relatively abundant. A good example is Lake Mohave, <br />AZ-NV, which supports the largest extant population of razorback sucker. The bonytail <br />presents a special challenge because it is not sufficiently abundant anywhere in nature <br />to afford opportunities for meaningful study. <br />Observations of behavior or habitat use for individuals at different stages of <br />development represent fragments of life history that must be assembled into a cohesive <br />story. The task is difficult because the subjects are hard to locate and there are <br />confounding factors, as mentioned previously. Nevertheless, a complete understanding <br />~ is important for the recovery effort because it can-help identify obstacles to recovery of <br />existing populations and sites with potential for restoring populations. The remainder of <br />the text in Part 2 of this report summarizes what is known about the life history <br />requirements of each of the four endangered fish species. <br /> <br /> <br />9 <br />• <br />