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Colorado squawfish. Hendrickson and Brooks (1987) noted predation by yellow <br />bullhead, Ameiurus natalis, and largemouth bass, Micropterus salmoides, on <br />young Colorado squawfish stocked into the Verde River, Arizona. Osmundson <br />(1987) noted predation by largemouth bass, green sunfish, black crappie, L. <br />nigromaculatus, and black bullhead, A. melas, on young Colorado squawfish in <br />gravel pits near the Colorado River and indicated that predation by channel <br />catfish may have occurred. In addition, Coon (1965) reported channel catfish <br />predation on Colorado squawfish in the Dolores River. <br />Humpback Chub <br />Humpback chub occur in several canyon-bound, turbulent sections of the <br />Green River basin, including the lower portion of the Little Snake River and <br />Yampa, Whirlpool, Desolation and Gray canyons (Holden 1978; Tyus et al. <br />1982ab, 1987; Karp and Tyus, in review; M. Moretti and E. Wick, pers. comm.). <br />Survey studies conducted prior to and following closure of Flaming Gorge Dam <br />indicated that all three forms of Colorado chubs were locally common in the <br />Green River; bonytails and roundtails in upper Green River above Ouray, Utah; <br />humpbacks in Desolation and Gray canyons and historically in Flaming Gorge <br />basin of upper Green River (Smith 1960; Holden 1978; Vanicek 1967; Seethaler <br />et al. 1979). Although intensive netting and electrofishing efforts in the <br />Green River mainstream by USFWS and Utah Division of Wildlife Resources <br />biologists yielded over 500 chubs (Miller et al: 1982; Tyus et al. 1987; M. <br />Moretti, pers. comm.), status of humpback chub in that system is not well <br />understood, due in part, to confusion with specific identification of some <br />individuals (Tyus et al. 1987; Rosenfeld and Wilkinson 1989; M. Moretti, pers. <br />comm.). <br />More intensive studies .are needed to further elucidate the systematics of <br />the Colorado River chubs and to determine their habitat needs in the Green <br />River basin. The morphological variation that occurs in some areas of sympatry <br />may be induced by recent habitat change (Valdez and Clemmer 1982). Thus, the <br />presence of intermediate forms in altered systems (e.g., Green River, Colorado <br />River) and the paucity of such forms in unaltered rivers (e.g., Yampa River, <br />Little Colorado River) suggests that natural riverine environments are <br />important for recovery of the humpback chub. The following paragraphs <br />summarize information on habitat use of humpback chub in Yampa Canyon because <br />this information is unknown for the Green River. We presume habitat use and <br />needs are similar throughout the Green River basin. <br />During the spring and early summer, adult humpback chub were most often <br />collected in shoreline eddy habitats created by large boulders and rapids in <br />Yampa Canyon (N=130; Karp and Tyus, in review). Three adult fish, one a ripe <br />male, were collected in Whirlpool Canyon. Juvenile fish were most prevalent in <br />smaller eddies in shoreline runs. Humpback chub were observed feeding at <br />various levels within the water column, consuming such terrestrial insects as <br />ants and Mormon crickets (Tyus and Minckley 1988; Karp and Tyus, in review). <br />There is some indication that humpback chubs in Yampa Canyon exhibit short- <br />distance, localized movements, and display some fidelity to specific eddy <br />habitats (Karp and Tyus, in review). <br />Adult humpback chub were captured in DNM in association with 18 other <br />fishes, but were most commonly captured with roundtail chub and the <br />introduced channel catfish when angling (Tyus and Minckley 1988; Karp and <br />Tyus, in review). Channel catfish comprised about 35% of the angling catch in <br />habitats from which humpback chub were collected (Karp and Tyus, in review) <br />21 <br />