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PART I: INTRODUCTION <br />General <br />The Colorado River drains about 244,000 mi' of the southwestern United <br />States and Mexico (Figure 1). Originating in the Never Summer and Wind River <br />ranges of Colorado, Utah, and Wyoming, it flows about 1,500 mi. through four <br />other western States (Arizona, California, Nevada and New Mexico) as it <br />travels toward its historic outfall into the Gulf of California in Mexico <br />(Carlson and Muth 1989). Most of the water in the system is snowmelt that <br />originates in high mountain streams generally above 10,000 ft amsl. Tributary <br />streams at lower elevations add comparatively little water. The natural <br />hydrograph reflects the regular and prominent influence of spring runoff in <br />May and June (Maddux et al. 1993; Stanford 1994). Peak flows produce <br />extensive seasonal flooding of overbank areas. Although the magnitude of the <br />peak runoff is variable, the timing of the event is predictable. Migh <br />discharge and erodible substrate produce very turbid water for much of the <br />year. Smaller tributaries are prone to flash flooding after unpredictable <br />summer storms. Storm events contribute to turbidity in the main river during <br />the base flow period. <br />The ancient Colorado River watershed was a much wetter environment than <br />now exists (Smith, M. 1981), and portions of the river system were dominated <br />by lacustrine habitat (Minckley et al. 1986). The basin is now much more <br />arid, and the river is more constrained within its banks, except for seasonal <br />flooding. Seasonal inundation is significant in part because wetland areas <br />1 <br />