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review) and the importance of this phenomenon needs further evaluation, <br />particularly in reintroduction efforts. <br />The capture and artificial spawning of ripe razorback suckers in the <br />lower Yampa and upper Green rivers (FWS unpublished data) and the tentative <br />identification of larvae in upper Green River seine collections (R. T. Muth <br />and D. Snyder, personal communication) indicates that razorback suckers <br />reproduce successfully in the upper Green River basin. However, there is <br />little indication of recruitment to the juvenile stage throughout the Colorado <br />River basin (Holden 1978; McAda and Wydowski 1980; Hinckley 1983; Tyus 1987; <br />Marsh and Hinckley 1989; Tyus and Karp, in review). Standing crops of <br />razorback suckers in the Green River are presumably old fish (Hinckley et al., <br />in prep) and no documented recruitment has occurred since the 1960s. <br />The apparent lack of widespread recruitment has been attributed to <br />habitat alteration, such as lower water temperatures (Marsh 1985) and <br />predation by introduced common carp Cyprinus carpio, green sunfish, channel <br />catfish, flathead catfish Pylodictus olivaris, and other non-native fishes- <br />(Minckley 1983; Brooks et a]. 1985; Tyus 1987; Marsh and Langhorst 1988;-Marsh <br />and Hinckley 1989). Predation by non-native fishes is believed a serious <br />threat to the survival and recovery efforts of razorback sucker. The absence <br />of young fish in the Green River basin population may-also be linked with the <br />reduced availability of inundated shorelines due to curtailment of spring <br />flooding following closure of Flaming Gorge Dam. Naturally inundated lowlands <br />such as Old Char]ey Wash and Stewart Lake Drain should be made accessible to <br />razorback suckers in the spring. The introduction of young fish (reared at <br />Ouray National Wildlife Refuge from sex products taken from wild fish captured <br />on nearby Green River spawning grounds) may ultimately result in natural <br />recruitment if the number of spawning adults, and consequently the number of <br />young fish, is increased. <br />Habitat requirements of this species in riverine environments are not <br />well known because of the scarcity of extant populations (Hinckley 1983; <br />Lanigan and Tyus 1989) and the absence of younger life.history stages <br />(McCarthy and Hinckley 1987; Tyus 1987j. Adult razorback suckers in the Green <br />River basin are old individuals (Tyus 1987, Hinckley et al ., in prep.) and the <br />low number of reproducing razorback suckers is considered limiting (Lanigan <br />and Tyus 1989; Tyus and Karp, in review). High spring flows, warming river <br />temperatures, and inundated shorelines and bottomlands are considered <br />essential to successful reproduction in this species. <br />PART III. SUMMARY AND RECOMMENDATIONS <br />Summary.: <br />General <br />The Green River basin is an important riverine system for the maintenance <br />and recovery of the endangered Colorado River fishes. Flows of the tributary <br />Yampa River are singularly important for maintaining a spring peak on the <br />Green River hydrograph (Figure 3), thereby mitigating some of the adverse <br />effects of flow regulation by Flaming Gorge Dam on the native fish fauna. <br />Flows of the Yampa. River, particularly spring runoff, may also enhance .usable <br />fish habitat by inhibiting the invasion-and proliferation of introduced fishes <br />that evolved in more mesic environments. Colorado squawfish, razorback sucker <br />and other fishes depend on habitats in the Yampa and Green rivers for <br />fulfillment of various life history requirements, and thus, the two rivers <br />must be viewed as a single system when determining flow needs. <br />24 <br />