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that covers all rivers at the same time, using the same methods (such as for the <br />Yampa, Green, White, Colorado,. Gunnison and Dolores Rivers in 1979-81: Miller et al. <br />1982ac, Tyus et al. 1982b, Valdez 1982), it was judged more acceptable to provide the <br />raw data than to artificially divide the database. -Thus, the capture data are used with <br />caution and they are supplemented with additional information as available. <br />The relative importance of tributary streams for supporting populations of each of the <br />listed species was evaluated using alife-history approach. This approach has both a <br />spatial and temporal component, because- life history stages may occupy different <br />habitats seasonally. Life history information was coupled with distributions of existing <br />populations in order to obtain a geographical perspective on potential roles for <br />tributaries. Next we assessed possible sites for establishing new populations, taking <br />into account the contributions that tributaries might make. <br />We determined the relative value of tributary streams as habitat for the endangered <br />fishes by assessing their direct habitat use, or potential for use above areas that are <br />blocked by barriers. Although native fish diversity is an important component of <br />endangered fish habitat, the presence of other native fishes was not a factor in our <br />ranking of tributary streams. The presence of other native fishes does not guarantee <br />that the four endangered fishes will occur sympatrically, and it remains to be proven <br />whether it will or will not be possible to recover the endangered fishes in the absence of <br />a native fish- community. However, the Endangered Species Act dictates that efforts be <br />made to conserve the ecosystem upon which listed species depend, and we provided <br />general information about native fish abundance to depict the present condition of the <br />streams in our evaluation. <br />We were unable to obtain much information about the relative amount of fish habitat <br />among tributaries, except as expressed in number of miles of river occupied by <br />endangered fishes. To obtain a crude basis for comparing tributaries, we obtained <br />measurements of habitat used by the endangered fishes from USGS topographic <br />maps. We obtained surface area and the volume of physical habitat in each tributary by <br />using a map wheel to measure the distance along the thalweg between adjacent <br />contour lines during the base flow period. Within each section thus delineated we <br />determined channel slope, width, and surface area. Using Manning's equation (Gordon <br />et al. 1992), we computed mean depth at base flow (median across years), assuming <br />that the roughness coefficient was 0.035 at all sites, and that the hydraulic radius is <br />essentially the same as depth for channels that are relatively wide. Mean depth was <br />then used to compute habitat volume and mean water velocity for each tributary reach <br />considered useful for one or more of the endangered fishes. <br />In addition to providing benefits to the endangered fishes, tributary streams also may <br />pose obstacles to recovery, due to a variety of factors. We iden#ified certain obstacles <br />when they were obvious, such as the presence of a barrier to fish movement, potential <br />water quality problems, presence of large numbers of nonnative fishes, and others. <br />However, we do not provide detailed analyses of these potential recovery obstacles. <br />7 <br />