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WATER PATHWAYS AND VECTORS <br />~`~~~ater pathways have probably been the most common means of spreading the zebra mussel in the United States. The mussels <br />are spread downstream as the veligers drift with the current. Once suitable habitats are reached, they attach to solid substrate <br />such as rock, piers, or boats. In upstream invasions, the veligers must be aided by a vector, which may include boats and <br />barges, or other carriers by which the mussel veliger can undergo passive transport. In contrast to land pathways, which are most <br />likely to be in straight lines, water pathways are not. As indicated in Figure 4, only two major types of water pathways exist: direct <br />conveyance of zebra mussels in lakes and rivers (downstream travel) and indirect transport by vessel. <br />$ASKAT~HEJrAN <br />\ , ~ MOt.~ANA <br />"^-.`~~ <br />~.. <br />ap ~-.,. Rive <br />~` <br />~ <br />~- ' <br />~ So ` <br />} <br />-_Y"_'_" ~L~ _. <br /> d~°ro„Q Rrur NOHTHOAKOTA ~~- <br /> _ ~ <br /> t <br /> WYOMING ~ <br />~ <br /> <br />~+ . <br />F» <br /> SOUTH DAKOTA <br /> Z~ <br />. <br /> 6 <br />_ .. " <br /> '+ NEBRASKA <br />P~/~ <br /> ~~ <br /> co~oaaoo ~.~~ <br />V ~ -. <br /> UTAH 'S~Ltb e <br /> ..~ KANSAS <br /> <br />+ <, AflIZONA ONUHOMA " <br /> , NEW ME%ICA - <br />', <br /> . <br />~°o <br />o . <br />o <br /> q gn <br /> RWR, <br /> ,,, <br /> <br /> <br />,.~' <br />Information on major river systems <br />in Europe which became infected with <br />the nonindigenous mussel has shown <br />that the mussel can be transported <br />great distances both upriver and <br />downriver within a short span of years. <br />Modes of its upstream transport are <br />generally not well understood, but <br />Carlton (1993) suggested that the zebra <br />mussel may be carried and transmitted <br />by a host of natural and man-influ- <br />enced means. All major rivers and <br />waterways entering and connecting the <br />western United States and eastern <br />infested areas should be considered <br />potential pathways (Figure 3). Major <br />waterways are identified in Figure 9. <br />F/GORE 9. MAJOR EAST-WEST R/VERB. <br />