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1 <br />1 <br />' calculated from larval size for 1981-1987. The length of optimal spawning <br /> period in the Yampa River (Table 2), derived from Figure 8, was similar for <br />' each year. Spawning generally occurred earl ier in low wa ter years (1981 and, <br /> 1987) and later with higher flows (1983 and 1984) (Table 2). Dates used in <br /> " <br />" <br />" <br />" <br /> estimation of <br />optimal <br />and <br />total <br />spawning periods are provided in Table 3. <br />' Total spawning period included any indication of spawning activity (e.g., <br /> radiotagged fish, collections of ripe fish, larvae) and was about 4 to 5 weeks <br />' in duration. Optimum spawning periods (indicating highest spawning activity <br /> as evidenced by simultaneous collections of radiotagged fish, ripe fish, and <br />' larvae) were somewhat shorter (Table 3). <br />' Table 3. Calculation of optimal spawning period for Colorado squawfish in <br /> <br />------ `Yampa River, <br />--- <br />- <br />- 1981-1987. <br />--- <br /> <br />' <br />Year - <br />- <br />------ <br />---- <br />Back 1 ----------- <br />Ripe -------------------- <br />Radiotagged ------------ <br />Spawning --------- <br />- <br />Period <br /> Calculations rFish FishZZ Total Optimum <br /> 1981 6/19-7/10 7/1-7/10 6/20-7/20 6/19-7/20 6/23-7/13 <br /> 1982 7/10-7/18 7/10-8/7 7/4-8/8 7/4-8/8 7/8-8/1 <br />' 1983 7/20-8/5 7/14-8/18 7/12-8/8 7/12-8/18 7/15-8/10 <br /> 1984 7/19-8/13 7/13-8/14 7/10-8/17 7/10-8/17 7/16-8/15 <br /> 1985 6/27-7/13 6/27-7/25 6/21-7/12 6/21-7/25 6/25-7/17 <br />' 1985 6/29-8/4 7/5-7/30 N/A 6/28-8/5 7/2-8/2 <br /> 1987 6/16-7/9 6/3-6/30 6/9-6/30 6/3-7/9 6/9-7/3 <br />' 1 Larvae were separated into two size classes: <22mm and 22-47mm, for back <br />calculation of larval age and estimation of spawning period (equations in <br />Haynes and Muth 1984, Kesler 1986). An estimate of total spawning period was <br />' obtained by combining estimated periods for two size classes. <br />2 Dates (range) represent first and last appearance of radiotagged fish on <br />spawning grounds. <br />Breeding adults were most often concentrated in river reaches containing <br />' deep pools and/or eddies and cobble (rubble) bars. The fish moved from pools <br />' and/or eddies to apparently spawn on cobble bars, then returned to the former <br />habitat (behavior similar to that reported for spawning northern squawfish <br />t (Beamsderfer and Congleton 1982)). Turbid conditions in the Yampa River <br />19 <br />