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<br />stable backwater conditions likely favored reproduction of the NNC, as well as enhanced the growth of <br />~ larvae into catchable subadult fish. Any reproduction fundamentally compromises closure assumptions <br />behind removal experiments at any spatial or temporal scale. <br />Lentsch et al. (1996) postulated that controlling nonnative cyprinids would be difficult because of <br />1 their high abundance, ubiquity, adaptability, and reproductive potential. Red shiner, fathead minnow, and <br />sand shiner are the most common species in the upper Colorado River basin, frequently comprising more <br />than 80% of fishes captured in low velocity habitats (Valdez 1990; Muth and Nester 1993; McAda et al. <br />1994a and 1994b, 1995, 1996, 1997; Trammell and Chart 1999a, 1999b, and 1999c). Minckley (1973), <br />Scott and Grossman (1973), Cross and Collins (1975), Pflieger (1975) Carlander (1977), Matthews <br />(1987), and Sublette et al. (1990) all report the tolerance of red shiner, sand shiner, and fathead minnow <br />~ to various perturbations of "quality" habitats or conditions. All three of these species are classified as <br />extremely tolerant of extremes in temperature, pollution, discharge, and siltation. Their small size and <br />ability to persist in and re-colonize a variety of habitats makes removal difficult. Cross and Collins (1975) <br />~ state that it will never be possible to completely remove red shiner and suggest they "will be with us until <br />the end, no matter how badly we treat our water". <br />If reduction in abundance was possible, detection of the reduction may also prove difficult. For <br />example, nonnative cyprinid data collected from 1986 to 1997 (McAda et al. 1994a, 1994b, 1995, 1996, <br />t <br />1997) during annual ISMP surveys in both the Green and Golorado rivers shows nonnative cyprinid <br />numbers vary greatly from year to year despite the use of standard sampling protocols (Figures 10, 18 <br />and 19). Furthermore, it appears that NNC density was at historically low levels in 1997, prior to removal <br />efforts. Some studies have shown that nonnative cyprinid abundance is reduced in high water years <br />(McAda et al. 1994a, McAda and Ryel 1998, Trammell et al. 1999, Lentsch et al. 1996); peak flows in <br />1997, 1998 and 1999 were all higher than average. Thus, due to inherent NNC population variability and <br />NNC density antecedent to this study, attributing any changes in NNC abundance to removal efforts is <br />difficult. <br />Fluctuating habitat availability (especially in the Colorado River) also contributed to difficulties in <br />interpreting data. Most of the variation in area sampled was due to differences in discharge and <br />-16- <br />