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19 <br />fluctuations in the backwaters resulted in a lower mean temperature than that of the main channel. <br />If Colorado squawfish follow temperature gradients closely, this would result in much greater <br />movement in and out of backwaters on a daily basis than previously surmised. <br />Temperature related to growth <br />Thompson (1991) reported that YOY squawfish cease growth at 13° C. Main channel and <br />backwater temperatures were greater than 13 from mid-March to mid-October. Since YOY <br />squawfish are first present in the river near the end of June, their fall growing season is about 3.5 to <br />4 months long. High peak flow was directly correlated to later hatch dates and lower degree day <br />accumulation (Trammell and Chart 1998; larval drift), and thereby to smaller total length of CS in <br />the fall (Figure 2). In low water years the fish spawned earlier, thereby lengthening the growing <br />season, and resulting in larger fish in the fall. Low water also allowed the river to warm up faster and <br />for backwaters to reach a higher maximum temperature. Degree day accumulation was calculated <br />from first hatch date to fall sample date, and regressed against fall TL, resulting in an rz of .65. When <br />1992 was removed from the regression, rz became .91. <br />The total length of squawfish in the fall was related to overwinter survival and continued overwinter <br />growth. The smaller the fish in the fall, the greater the decline in catch rates from fall to spring <br />(Figure 3), which suggests some size-related overwinter mortality. However, all years except `95 <br />showed some growth occurring between the fall sampling and the spring sampling. Maximum, <br />minimum and average TL, all increased over winter (Table 9). Fall and spring sampling dates <br />bracketed the dates the river fell below and rose above 13° C, with a few days above 13° C on either <br />end. <br />