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<br />Stanford [ 1994] stated that the Andrews and Nelson 1989 model was otentiall an a ro riate ' <br />[ ] p Y pp p <br />addition to the Upper Colorado basin research program. <br />Flow Re mm ' <br />co endataon Research <br />Historically, instream flow recommendations have been based on the flows necessary for ' <br />maintaining a suitable amount of fish habitat during base flows [e.g., Tennant, 1975]. More <br />recent work suggests that a suite of flows is necessary to maintain ecological integrity offish ' <br />habitat rather than just protecting the habitat at base flows [Hill et al., 1991 ]. This latter <br />approach is based on the assumption that habitat and nutrient availability for the fishes require <br />maintenance of the linkages between the stream and riparian, floodplain and upland areas. ' <br />Two philosophies have shaped the design of instream flow for Colorado River native <br />fishes [Tyus, 1992]. The first philosophy is to provide minimum flows needed for species <br />survival during periods of low river discharge. These minimum flows do not ensure availability ' <br />of optimal habitat, but seek to maintain and maximize the availability of existing habitat. The <br />second philosophy is to assist species recovery by increasing the availability of optimal habitats, <br />requiring that existing habitat be improved, and restoring "lost" habitats (i.e., habitats that are no <br />' <br />longer formed or available as a consequence of flow regulation). "Optimal" habitats under <br />regulated conditions include conditions of altered river hydrology and the presence of exotic <br /> <br />species, and are not the pristine river conditions that existed prior to regulation. For example, a ' <br />habitat that was optimal prior to river regulation may no longer be optimal due to its use by <br />predatory species. For species recovery to occur in a system as altered, both physically and ' <br />biologically, as the Green River, the second philosophy, with its emphasis on optimal habitats <br />and increased habitat availability, is the only viable option for species recovery. <br /> ' <br />Multiscale, Multiflow Studies <br />Ligon et al. [1995] noted the need for multiscale, interdisciplinary studies to quantify and <br />mitigate the downstream effects of dams. Hill et al. [1991] integrated several standard ' <br />assessment methods into a methodology for determining the suite of flows necessary to maintain <br />ecological integrity and applied their methodology to the Salmon River at Whitebird, Idaho. <br />They noted the need for four flow components: base flows for fish, and channel-, riparian-, and <br />' <br />valley-process maintenance flows. Each of these flows inundated different ecosystems (e.g., <br />stream channel and riparian ecosystem), and combinations of methodologies were necessary to <br /> <br />evaluate the appropriate flow regime for each component. ' <br />Base Flows <br />Habitat availability curves were first developed for cold headwater streams to assess the ' <br />effects of water diversions on standing crops offish [Nesler, 1990]. The US Fish and Wildlife <br />Service's Instream Flow Incremental Methodology (IF'IM) and Physical Habitat Simulation <br />system (PHABSIM) [Bovee, 1982] are widely used to predict discharge-dependent changes in ' <br />area of available habitat. PHABSIM uses suitability curves determined from measurements of <br />physical parameters to model weighted usable area (WUA), a surrogate for the area of available <br />habitat for a target species. This model assumes that a positive, linear relationship exists , <br />between WUA and fish biomass, which implies that the target fish population is habitat limited <br />[Mathur et al., 1985]. <br />PHABSIM [Bovee <br />1982] was used b <br />t al <br />[1991] t <br />uantif <br />low <br />Hill <br />th <br />t <br />fl <br />h <br />bit <br />' <br />, <br />- <br />y <br />e <br />. <br />o q <br />y <br />e <br />ow <br />a <br />a <br />A-10 , <br />