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u <br />1 <br />1 <br /> <br /> <br /> <br />1 <br /> <br /> <br /> <br />1 <br /> <br /> <br />1 <br /> <br /> <br /> <br />1 <br />1980, Hinckley et al. 1986}. Hinckley et al. (1986} divided freshwater fishes of the CRB <br />into three major categories. The first group, big-river fishes, included largely endemic <br />species that occur in large streams and rivers. The second group was described as <br />endemic fishes occurring in small streams at low to intermediate elevations. The final <br />group included fishes occupying mid- to high-elevation habitats with relatives co- <br />occurring or present in adjacent drainages outside the CRB. Although also found in <br />small streams, roundtail chub, flannelmouth sucker, and bluehead sucker can be classifed <br />in the first group of mostly endemic, big-river fishes. <br />Several morphological characteristics are common to the native, big-river fishes, <br />including streamlined body forms, humped or keeled dorsal surfaces, leathery skins with <br />fine or embedded scales, and large, often falcate, fins. These characteristics provide for <br />improved swimming performance and stability in turbulent flows, which are common in <br />much of the Colorado River and tributaries (Hubbs 1941, Hinckley et al. 1986, Hinckley <br />1991). Such morphologic features, combined with comparatively long life spans and <br />breeding seasons initiated by water temperature and flow regime cues, may be <br />adaptations to the harsh, unpredictable physical environment of CRB rivers and streams <br />(Scoppettone 1988, Hinckley 1991, Stearns 1993). Prior to Euro-American settlement, <br />the suite of native, big-river fishes present in the CRB were well-adapted to the <br />environmental extremes of the basin, as several taxa have persisted for over 10 million <br />years (Hinckley et al. 1986). <br />Human Impacts <br />If declines in big-river fishes over the past half century are any indication, human-caused <br />changes have reduced the effectiveness of morphologic and life-history adaptations that <br />allowed native fishes to thrive in the historically harsh, fluctuating environment of the <br />CRB. In the late 1800s, settlers moving into the American Southwest demanded <br />legislation to fund major water development projects. Since then, the water projects <br />authorized by these acts and legislation (Table 2), and the agriculture, infrastructure, and <br />Final Report September 2002 <br /> <br />