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Acknowledgements <br />We thank R. J. Behnke, K. R. Bestgen, C. A. Carlson, K. D. Fausch, <br />J. A. Hawkins, D. E. Snyder, R. A. Valdez, and J. A. Wiens, for comments that <br />improved the manuscript. We are grateful to P. J. Sikoski, and J. M. Bundy <br />for assistance in the laboratory. This study was funded by the Recovery <br />Implementation Program for Endangered Fish Species in the Upper Colorado River <br />Basin. This Program is a joint effort of the U. S. Fish and Wildlife Service, <br />U. S. Bureau of Reclamation, Western Area Power Administration, States of <br />Colorado, Utah, and Wyoming, Upper Basin water users, and environmental <br />organizations. <br />24 <br />