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<br />Importance of Tributaries ' <br />Tributaries of the Green and Colorado Rivers may be especially important to <br />the continued survival of the Colorado squawfish. Colorado squawfish ' <br />historically or presently occupy tributaries of the Green and Colorado Rivers, <br />including the Yampa, Duchesne, White, Gunnison, Dolores, and San Juan Rivers <br />in the-Upper Basin, the Gila River and its tributaries, the Salt, Verde, and , <br />other rivers in the Lower Basin. Tagging and telemetry studies indicate- the <br />Colorado squawfish moves long distances from spawning areas in the mainstem <br />Green River to feeding and overwintering areas in the Yampa, White, and <br /> <br />Duchesne Rivers (Miller et al. 1982a, 1982b, 1983; Radant et al. 1983; Tyus et ' <br />al. 1982a, 1982b, 1987). Tyus (1986) suggested that tributaries play an <br />important role in the life history strategy of the Colorado squawfish and <br />perhaps serve to reduce intraspecific competition. ' <br />Miller et al. (1982a) reported a net movement of juveniles from the Green <br />River to the White River, while Radant et al. (1983) documented movement of <br /> <br />adult fish between the White and Green Rivers and Green and Yampa Rivers ' <br />during the spawning season. There also is a downstream movement of larval <br />Colorado squawfish from the Yampa River to the Green River (Haynes et al. <br />1984; Miller et al. 1982b; Tyus and Haines 1991). Furthermore, a net upstream <br />movement of adult and juvenile Colorado squawfish from the Green River into <br />the Yampa River has been suggested by Miller et al. (1982b) and Tyus (]986, <br />1990). <br />Colorado squawfish larvae have been collected from the Yampa River within <br />Dinosaur National Monument each year during the period 1980-88 (Nesler et al. <br />1988; Tyus and Haines 1991; Wick et al. 1981), and spawning migrations into <br />the lower Yampa were observed in 1981-88 (Wick et al. 1983; Tyus 1990). <br />Holden (1980) and Tyus and Karp (1989) indicated that flows from the Yampa <br />River were important to the recruitment of Colorado squawfish in the Green <br />River. Analysis of the hydrographs indicates that the Yampa River is the <br />primary contributor of high spring flows in the Green River which generally <br />exceeded 12,000 cfs at the Jensen gage. Spring flows of the Yampa River <br />affect the timing of Colorado squawfish spawning migrations (Tyus 1990). <br />Reasons for Decline ' <br />Historically, the Colorado River was characterized as a river with wide <br />seasonal flow fluctuations (Waters 1946). It was known as Rio Colorado, the <br />great Red River of the West. Over 2,700 km (1,687 miles) long, it is cliff ' <br />bound in canyons throughout much of its course. It drops over 3.2 km <br />(2 miles) in its journey to the Gulf of California, thereby creating some of <br />the most turbulent waters found on earth. At Lee's Ferry, Colorado River ' <br />streamflows varied from 750 cfs in 1924 to an estimated 300,000 cfs in 1884 <br />(White and Garrett 1988). Few rivers were so laden with silt. Averaging <br />0.62 percent silt content by volume, it formerly carried more than ' <br />100,000 acre-feet (12,000 ha-meter) of soil to the Gulf of California each <br />year. It also is high in mineral salts: carbonates; sulfates; and chlorides <br />of calcium, sodium, and magnesium. <br /> <br />.. <br /> ' <br />