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<br />Life History , <br />Habitat Preference <br />h <br />hi <br />fl <br />- <br />s ' <br />ow, <br />g <br />easonally variable <br />Colorado squawfish <br />are adapted to rivers with <br />silt loads, and turbulence. Young-of-the-year (up to 64 mm [2.5 in.] TL), <br />juveniles (65-200 mm [2.5-8 in.]), and subadults (200-400 mm [8-16 in.]) live <br />in shallow backwater areas, with little or no current over silt and sand <br />bottoms (Haines and Tyus 1990; Holden 1913; Holden and Stalnaker 1975a, 1975b; <br />Holden and Twedt 1980; Miller et al. 1982a, 1982b; Tyus and Haines 1991; <br />Ualdez et al. 1982b; Valdez and Wick 1983; Wick et al. 1979, 1981). There is <br />a change in habitat preference at about 200 mm (8 in.) TL (Miller et al. <br />1982a), with larger fish selecting deeper water of at least some velocity. <br />Adults are large-river fish, found in a variety of depths and velocities over , <br />silt, sand, gravel, and boulder substrates (Holden 1973; Holden and Twedt <br />1980; Holden and Wick 1982; Miller et al. 1982a, 1982b; Tyus et al. 1982a, <br />1984; Valdez et al. 1982b; Wick et al. 1979, 1981). ' <br />Radiotelemetry studies (Miller et al. 1983; Tyus 1985, 1986; Tyus and McAda <br />1984; Tyus et al. 1982b; Wick et al. 1983) have provided considerable <br />information on habitat usage of adult Colorado squawfish. Adults use various ' <br />habitats depending upon season, streamflow, water temperature, and <br />availability (Holden and Wick 1982; Tyus and McAda 1984; Tyus 1990; Wick et <br />al. 1983, 1985, 1986). During peak runoff, adults move into backwater areas <br /> <br />or flooded riparian areas where velocity is lower and water temperatures are ' <br />higher than in the main channel (Wick et al. 1983). During the decline in <br />water level following peak runoff, spawning adult fish move into run-riffle <br />areas and also occupy run, eddy, and pool habitats (Tyus 1990). Adult <br />Colorado squawfish exhibited little movement during winter (October-April) in <br />the upper Green River (Valdez and Masslich 1989). Of 20 adults radio-tagged <br />in October, 15 moved less than 5 km by the end of the following March. The ' <br />fish occupied primarily slow runs, slackwater, eddies, and backwaters. <br />Temperature Preference ' <br />The thermal tolerance of Colorado squawfish is broad, as evidenced by the <br />range of temperatures to which the species was presumably adapted. Summer ' <br />water temperatures in the vicinity of Yuma, Arizona, for example, commonly <br />approach or exceed 35oC (95oF) and may drop to lower than lOoC (50oF) in <br />winter (Minckley 1979). In the Upper Basin, water temperatures generally <br />range from 25oC (77oF) during the summer to freezing (OoC [32oF]) in winter. ' <br />Tyus (1990) summarized the water temperatures associated with prespawning, <br />migratory, and spawning periods for adult Colorado squawfish in the Upper <br />Green River Basin. Spawning migrations were initiated at water temperatures <br />of 14-20oC (57o-68oF), and spawning occurred at temperatures of 22oC (72oF) <br />(range 15-27.5C [59o-82oF]). In the Yampa River, migrations and spawning <br />periods varied between years. Migrations were initiated from May 12, to <br />June 10, associated with a mean water temperature of about 14oC (57oF), and <br />spawning occurred at 21oC (70oF) (Table 1) (Tyus and Karp 19$9). However, <br />Tyus et al. (1987) and Wick et al. (1985) cautioned that main channel ' <br /> <br /> <br />