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27 <br />(Sigler and Miller 1963), Desolation Canyon of the Green River (Holden and <br />Stalnaker 1970), and the Colorado River near Moab (Sigler and Miller 1963). <br />Today the largest populations of this species occur in the Little Colorado <br />River, in the Grand Canyon, and in the Black Rocks area of the Colorado River. <br />Other populations have been reported in Westwater and Debeque Canyons~of the <br />Colorado River, Desolation and Gray Canyons of the Green River, Yampa and <br />Whirlpool Canyons in Dinosaur National Monument (USFWS 1990b). One individual <br />was recently captured in the Gunnison River (Burdick 1995). <br />Little is known about the specific spawning requirements of the humpback chub. <br />The fish is known to spawn soon after the highest spring flows when water <br />temperatures approach 68° F (Kaeding. et al. 1990. Karp and Tyus 1990a, USFWS <br />1990b). The collection of ripe and spent fish indicated that spawning <br />occurred in Black Rocks during June 2-15, 1980, at water temperatures of 53° <br />to 61° F; in 1981, spawning occurred on May 15-25 at water temperatures of 61° <br />to 61.7° F (Valdez et al. 1982b). Humpback chub spawned in Black Rocks on the <br />Colorado River in 1983 when maximum daily water temperatures were 55° to 63° F <br />(Archer et al. 1985). <br />Backwaters, eddies, and runs have been reported as common capture locations <br />for young-of-year ~~umpback chub (Valdez and Clemmer 1982). These data <br />indicate that in Black Rocks and Westwater Canyon, young utilize shallow <br />areas. Habitat suitability index curves developed by Valdez et al. (1990) <br />indicate young-of-year prefer average depths of 2.1 feet with a maximum of 5.1 <br />feet. Average velocities were reported at 0.2 feet per second. <br />Valdez et al. (1982b) and Wick et al. (1979, 1981) found adult humpback chub <br />in Black Rocks and Westwater Canyons in water averaging 50 feet in depth with <br />a maximum depth of 92 feet. In these localities, humpback chub were <br />associated with large boulders and steep cliffs. <br />Generally, humpback chub show fidelity for canyon reaches and move very little <br />(Miller et al. 1982c, Archer et al. 1985; Burdick and Kaeding 1985, <br />Kaeding et al: 1990). Movements of adult humpback chub in Black Rocks on the <br />Colorado River were essentially restricted to a 1-mile reach. These results <br />were based on the recapture of Carlin-tagged fish and radiotelemetry studies <br />conducted from 1979 to 1981 (Valdez et al. 1982b) and 1983 to 1985 (Archer et <br />al. 1985; USFWS 1986; Kaeding et al. 1990). <br />