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25 <br />Specific information on biological and physical micro-habitat requirements of <br />the razorback sucker is very limited. Until very recently, fisheries research <br />investigations throughout the Upper Basin have focused on the other three <br />listed Colorado River fishes, and data on the razorback sucker were collected <br />largely coincident to those studies. Localized extirpation of razorback <br />suckers from some localities, coupled with the species' continued decline in <br />numbers and distribution, has prompted some research; however, details of its <br />life history requirements, particularly in riverine environments, are still <br />not fully understood. <br />In general, a natural hydrograph with a large spring peak, a gradually <br />descending limb into early summer, and low stable flows through summer, fall, <br />and winter are thought to create the best habitat conditions for endangered <br />fishes while maintaining the integrity of the channel geomorphology. Prior to <br />construction of large mainstem dams and the suppression of spring peak flows, <br />low velocity, off-channel habitats (seasonally flooded bottomlands and <br />shorelines) were commonly available throughout the Upper Basin (Tyus and Kar~~ <br />1989; Osmundson and Kaeding 1991). The absence of these seasonally flooded <br />riverine habitats is believed to be a limiting factor in the successful <br />recruitment of razorback suckers in their native environment (Tyus and Karp <br />1989, Osmundson and Kaeding 1991). Tyus (1987) and McAda and Wydoski (1980) <br />reported springtime aggregations of razorback suckers in off-channel <br />impoundments and tributaries; such aggregations are believed to be associated <br />with reproductive activities. Tyus and Karp (1990) and Osmundson and Kaeding <br />(1991) reported off-channel habitats to be much warmer than the mainstem river <br />and that razorback suckers presumably moved to these areas for feeding, <br />resting, sexual maturation, spawning, and other activities associated with <br />their reproductive cycle. While razorback suckers have never been directly <br />observed spawning in turbid riverine environments within the Upper Basin. <br />captures of ripe specimens, both males and females, have been recorded (Valdez <br />et al. 1982, McAda and Wydoski 1980, Tyus 1987, Osmundson and Kaeding 1989, <br />Tyus and Karp 1989, Tyus and Karp 1990, Osmundson and Kaeding 1991,;Platania <br />1990) in the Yampa, Green, Colorado, and San Juan rivers. Sexually mature <br />razorback suckers are generally collected on the ascending limb of the <br />hydrograph from mid-April through June and are associated with coarse gravel <br />substrates (depending on the specific location). <br />Outside of the spawning season, adult razorback suckers occupy a variety of <br />shoreline and main channel habitats including slow runs, shallow to deep <br />pools, backwaters, eddies, and other relatively slow velocity areas associated <br />with sand substrates (Tyus 1987, Tyus and Karp 1989, Osmundson and Kaeding <br />1989, Valdez and Masslich 1989. Osmundson and Kaeding 1991, Tyus and Karp <br />1990). <br />