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258 C. A. KARP AND H. M. TYUS [Volume 50 <br />UTAH ; COLORADO ~- ~ <br /> ARE <br />E,y <br /> <br />~~L <br />F9 ~ <br /> <br /> <br />u r a <br />~~ I <br /> <br />0 <br /> ~P ~ <br />~ (vA^~~ <br />1 y <br />` 0 <br /> P <br />~~ ~ _I <br /> SAP <br /> <br /> <br />~ VE <br />4S YQ A/ <br />Pq <br />p~nnaaar <br /> ool <br />~~,d <br />un Nac~onai p~yFq CRALG <br /> ?~~~ <br />,la~^ a ~anY Mon umenl <br />M AY B E L L• ~ <br /> Gmi Yampa ~anYOn <br />p,Q <br />P, Echa <br />Parts ~ <br />peerlotlge e <br />Park M'ILLI ,ā€ž <br />A S <br />Gpb~~ <br />SCaIE Ix MILES <br />r5 <br />aULE IH rNILOMEiEPS FpgK <br />Fig. 1. Yampa and Green rivers, Colorado and Utah, showing the boundaries of the study area and Dinosaur <br />National Monument. <br />at least twice each spring in 1987 and 1988, <br />and Lodore Canyon (Green River) was sam- <br />pled once each spring. Survey sampling (in- <br />cluding trammel netting) was conducted <br />throughout the Monument in July 1986 to <br />locate humpback chubs. Use of trammel nets <br />was discontinued after this effort because of <br />trauma to the fish. <br />Sampling trips in Yampa Canyon occurred <br />at weekly intervals (preceding and following <br />first and last capture of ripe fish) to insure an <br />accurate assessment of the humpback chub <br />spawning period. Our efforts were less inten- <br />sive in the Green River portion of the Monu- <br />ment because earlier sampling had yielded <br />few adult chubs in these reaches (Holden and <br />Crist 1981, Miller et al. 1982). Sampling pre- <br />ceded peak flows and was suspended during <br />highest runoff (2-4 week period depending on <br />water year) because of sampling problems in <br />high water. Sampling ended each summer <br />with attainment of base flows (approximately <br />late June to early July). Our efforts were re- <br />stricted to the spring and early summer be- <br />cause of boat accessibility. However, two <br />areas in Yampa Canyon that yielded hump- <br />back chub in the spring (Big Joe Rapid and <br />vicinity, Warm Springs Rapid and vicinity) <br />were sampled in September 1989 via heli- <br />copter and by foot to assess habitat, availabil- <br />ity, use, and substrate composition during <br />low flows. <br />All chubs greater than 85 mm total length <br />(TL) were identified to species using estab- <br />lished morphological characters (Smith et al. <br />1979, Douglas et al. 1989). We did not evalu- <br />ate habitat use of young humpback chub be- <br />cause we could not reliably distinguish young <br />of the various Gila species. Humpback chub <br />greater than 250 mm TL were tagged with <br />uniquely numbered Carlin-dangler tags for <br />recapture information (e. g. , growth and move- <br />ment data). Sex determination was based only <br />on expression of eggs or milt from ripe fish, <br />either spontaneously or following manual <br />pressure on the abdomen. Fish with breeding <br />tubercles but without expressible sex prod- <br />ucts were considered in reproductive condi- <br />tion. <br />Riffles, small rapids, runs, eddies, pools, <br />and backwaters were sampled. Because wa- <br />ter turbidity precluded visual contact with <br />humpback chub, it was necessary to esti- <br />mate the point of capture. Physical habitat <br />parameters recorded at each humpback chub <br />capture included water depth, temperature, <br />and substrate type. Depth was measured <br />with a calibrated rod, gross substrate type was <br />described from visual and manual examina- <br />tion, and temperatures were obtained with <br /> <br /> <br /> <br />