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Wyoming <br />N " <br /> Yampa <br /> ~i <br /> Q. White <br />0 100 200 <br /> ~ A <br />Kilometers ~ Colorado <br /> ~ <br />Utah <br /> Gunnison <br />Nevada <br />et <br />J <br />Cp~o <br />Q;~ <br />rado <br /> San Juan <br />B <br /> New Mexico <br />Arizona <br />California <br />C <br />Gila River <br /> Mexico <br />Figure 1. Map of the Colorado Hiver basin showing rivers that historically" <br />provided habitat for Colorado sgauwfish (light lines) and those <br />that presently contain Colorado squawfish (heavy lines). (Modified <br />from Seethaler 1978.) Sites of temperature-data collection in <br />upper (A), middle (ii) and lower (C) river reaches are indicated. <br />Seethaler (1978) examined the gonads of 147 upper-basin Colorado <br />squawfish 184--652 mm long and concluded that the onset of sexual maturity <br />occurred about 428 mm TL (6 years of age) (Figure 2). Spawning of Colorado <br />squawfish in the upper basin normally begins in July or August, when water <br />temperatures reach 20-22 C. This conclusion is based on probable spawning <br />activity of radiot"egged adult. Colorado squawfish (Tyus and McAda 1984), on the <br />subsequent occurrence of larvae (Haynes et al. 1984), and on observations on <br />the spawning activity of captive adults (Hammon 1981). Additional information <br />on the life history of Colorado squawfish was provided by Seethaler (197$) and <br />Holden and Wick (1982). <br />The Colorado squawfish of the upper basin is relatively slow--growing and <br />late-maturing--life-history characteristics unusual to be those of a principal <br />piscavore. Normally, successful predators grow rapidly (to better use prey <br />148 <br />~, ., .~..~,....__m~,...~. ~..,~.,,~,A. <br />