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588 <br />4 <br />rn <br />3 <br />w <br />m <br />Y <br />m <br />2 <br />U <br />c <br /> <br />1 <br />0 <br />KAEllING AND ZSMMERMAN <br />0 <br />c Clear Water <br />* Turbid Water <br />0 <br />0 <br />0 <br />0 <br />0 <br />0 <br />.o <br />• Minnow Trap <br />O Seine <br />• • <br />O <br />,6 <br />•2 • <br />.4 <br />• • <br />O <br />O <br />.2 <br />p •2 <br />•2 <br />• O o <br />0 5. 04 • 82 02 2. 002 <br />a <br />F <br />3 <br />°e <br />C_ <br />U <br />c <br /> <br />4 <br />3 <br />c <br />y <br />m <br />Y <br />2 m <br />O <br />U <br />c <br />t <br />0 <br />Mean Clf -Daylight Seining LC 2 LC 3 LC 4 C 4 <br />Little Colorado River Confluence <br />FtcuxE 7.-Relationships between mean catch rates by~ Flcuxe 8.-t1~leaaz catch rates for age-0 and juvenile <br />seine during darkness (2-14 seine hauls per mean; aver- humpback chubs collected by minnow traps and seines <br />age 6.9) and during d¢ylight (Z-18; 9.8) far age-0 and during darkness from reaches LC 2, 3, and d of the Little <br />juvenile humpback daubs in the Little Colorado River, <br />confluence, and Colorado River, 1980-181. Each data Colorado River and from tFee confluence of this tributary <br />point gives the mean catch rates for the 2-day collection wath the Colorado River, 1980-1981. Numbers desig- <br />nate multiple observations. <br />effort at a sampling site; efforts that yielded no humpback <br />chubs in both diet periods are not included. Seven data <br />paints are included in the aggregation near the origin. <br />humpback chubs avoid the clear, shallow water <br />swept by our seines during daylight, a hypoth- <br />esis supported by the occurrence of humpback <br />chubs in these same littoral areas when the water <br />is turbid. <br />Mean C/f for age-0 humpback chubs collect- <br />ed byminnow trap and by seine during darkness <br />from the Little Colorado (data largely from <br />1981) increased with proximity to the conflu- <br />ence (Fig. 8). Although this relationship sug- <br />gests that downstream reaches were most im- <br />portant for humpback chub reproduction in <br />1981, downstream drift or movement of small <br />fish could account for this distribution. Data on <br />seining during daylight for the postspawning <br />period in 1980 and 1981 are more detailed than <br />are the data for seining during darkness, and <br />they provide evidence of differences in hump- <br />back chub reproductive success between years. <br />Mean C/f for age-0 humpback chubs was ap- <br />preciably higher in 1980 than in 1981; density <br />was greatest at LC 2, where no age-0 humpback <br />chubs were collected in 1981 (Table 4). Sea- <br />sonalrunoff coincident with spawning could sig- <br />nificantly enhance humpback chub reproduc- <br />tive success in the Little Colorado, particularly <br />in upstream regions. Seasonal runoff occurred <br />during the spawning season in 1980 but not in <br />1981. Humpback chubs that spawned in hatch- <br />ery raceways laid eggs that adhered to rock sub- <br />strate where they were deposited (Hammon <br />1982). Runoff of the Little Colorado removes <br />fine travertine sediments that could suffocate <br />developing humpback chub embryos. Runoff <br />also dilutes chemically concentrated spring <br />waters, which otherwise might be harmful to <br />embryos and larvae. <br />Adults <br />Of 504 humpback chubs averaging 2'78 ± 54 <br />(SD) mm long collected in trammel nets, 93% <br />were classified as adults. Higher catch rates dur- <br />ing sunset and darkness than during daylight <br />suggested that activity of the fish increased as <br />light diminished, although avoidance of nets <br />during daylight cannot be discounted. Many of <br />