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~-. 800 <br />700 <br />H 600 <br />Z <br />W 5~ <br />J <br />J 400 <br />H <br />300 <br />Q zoo <br />~ ioo <br />Northern pike I <br />Largemouth bas j~ <br />I ~-• <br />1 ~- <br />.~ I <br />~' I Colorado squawfish <br />,' I ~ (River) <br />'I / <br />'~`._ q <br />Colorado s uawfish <br />(Pond) <br />Z ~ 4 5 6 7 8 9 10 11 <br />YEAR OF LIFE <br />FIGURE 2. Comparison of growth rate of Colorado squawfish from the <br />upper Colorado River basin (Vanicek and Kramer 1969, Seethaler <br />1978) with these characters for northern pike and largemouth <br />bass (vertical bars are the median 50 percent of the values <br />provided for these latter species by Carlander 1969, 1977). <br />Also shown is growth of Colorado squawfish in a pond near <br />Grand Junction, Colorado (Osmundson, 1986). The dashed line <br />on the curve for pond fish is projected growth. <br />adjacent upper Colorado River, the short growing seasons provided <br />by upper basin rivers were implicated as the cause of slow growth <br />of Colorado squawfish therein. <br />Growing season for Colorado squawfish doubtless was longer in <br />downstream regions of former range, but how rapid growth there <br />might have been is unknown because squawfish were extirpated from <br />these areas before such life-history information could be <br />collected. An understanding of the relative length of growing <br />seasons provided by these former habitats can be obtained, however. <br />Black and Bulkley (1985) studied the relation between temperature <br />and growth of Colorado squawfish with excess food. Our least- <br />squares analysis of their Table 1 data suggested no growth will <br />occur below about 14 C. If we assume the growing season is when <br />water temperatures exceed 14 C, that of the present-day upper basin <br />Colorado River is less than five months whereas those of the <br />historic middle and lower river were about six and eight months, <br />respectively (Figure 3). Similarly, the Osmundson (1986) pond <br />-110- <br />