c
<br />;i
<br />ICHTHYOLOGICAL NOTES
<br />formerly might have included bonytail habitat.
<br />Water temperatures there are most affected by
<br />air temperature; discharge is the fifth most im-
<br />portant independent variable affecting water
<br />temperature (after air temperature, solar ra-
<br />diation, wind speed and cloud cover, respec-
<br />tively) (Bob Green, Regional Hydrologist, U.S.
<br />Fish and Wildlife Service, Denver, Colorado,
<br />pers. com.). Nonetheless, bonytail are very rare
<br />in these areas. The same holds true for the low-
<br />er Green, where temperatures are not affected
<br />by operation of Flaming Gorge Dam but where
<br />a viable bonytail population no longer occurs.
<br />Factors other than alteration of temperature
<br />regimes must have eliminated bonytail from
<br />these river reaches. This same conclusion was
<br />reached by Marsh (1985) for some reaches of
<br />the lower Colorado. Operation of upstream
<br />dams has reduced seasonal-high discharges and
<br />increased seasonal-low discharges in these
<br />reaches, but it seems unlikely that a species like
<br />bonytail-which evolved under the wide vari-
<br />ety of discharge conditions characteristic of the
<br />seasonally dynamic Golorado River system-
<br />would have been affected directly by such mod-
<br />eration of discharge regimes. These river
<br />reaches have been successfully colonized by sev-
<br />eral introduced species (Tyus et al., 1982); per-
<br />haps these species or synergic effects of these
<br />species and the altered discharge regimes have
<br />had important effects on bonytail.
<br />Acknowledgments.-We thank G. H. Clemmer,
<br />P. B. Holden, R. R. Miller, W. L. Hinckley and
<br />G. R. Smith for their opinions on the identity
<br />of the bonytail. We thank colleagues and an
<br />anonymous reviewer for criticisms that greatly
<br />improved this manuscript. These observations
<br />were made as part of an investigation funded
<br />by the U.S. Bureau of Reclamation and the Mu-
<br />nicipal Subdistrict, Northern Colorado Water
<br />Conservancy District.
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<br />
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