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c <br />;i <br />ICHTHYOLOGICAL NOTES <br />formerly might have included bonytail habitat. <br />Water temperatures there are most affected by <br />air temperature; discharge is the fifth most im- <br />portant independent variable affecting water <br />temperature (after air temperature, solar ra- <br />diation, wind speed and cloud cover, respec- <br />tively) (Bob Green, Regional Hydrologist, U.S. <br />Fish and Wildlife Service, Denver, Colorado, <br />pers. com.). Nonetheless, bonytail are very rare <br />in these areas. The same holds true for the low- <br />er Green, where temperatures are not affected <br />by operation of Flaming Gorge Dam but where <br />a viable bonytail population no longer occurs. <br />Factors other than alteration of temperature <br />regimes must have eliminated bonytail from <br />these river reaches. This same conclusion was <br />reached by Marsh (1985) for some reaches of <br />the lower Colorado. Operation of upstream <br />dams has reduced seasonal-high discharges and <br />increased seasonal-low discharges in these <br />reaches, but it seems unlikely that a species like <br />bonytail-which evolved under the wide vari- <br />ety of discharge conditions characteristic of the <br />seasonally dynamic Golorado River system- <br />would have been affected directly by such mod- <br />eration of discharge regimes. These river <br />reaches have been successfully colonized by sev- <br />eral introduced species (Tyus et al., 1982); per- <br />haps these species or synergic effects of these <br />species and the altered discharge regimes have <br />had important effects on bonytail. <br />Acknowledgments.-We thank G. H. Clemmer, <br />P. B. Holden, R. R. Miller, W. L. Hinckley and <br />G. R. Smith for their opinions on the identity <br />of the bonytail. We thank colleagues and an <br />anonymous reviewer for criticisms that greatly <br />improved this manuscript. These observations <br />were made as part of an investigation funded <br />by the U.S. Bureau of Reclamation and the Mu- <br />nicipal Subdistrict, Northern Colorado Water <br />Conservancy District. <br />LITERATURE CITED <br />BOZEK, M. A., L.J. PAULSON AND J. E. DEACON. 1984. <br />Factors affecting reproductive success of bonytail <br />chubs and razorback suckers in Lake Mohave. Lake <br />Mead Limnological Res. Cent., Dept. Biol. Sci., <br />Univ. Nevada, Las Vegas. Tech. Kept. No. 12. <br />GILBERT, G. H., AND N. B. SGOFIELD. 1898. NOteS On <br />a collection of fishes from the Colorado basin in <br />Arizona. Proc. U.S. Natl. Mus. 20:487-499. <br />HAMMAN, R. L. 1981. Hybridization of three species <br />of chub in a hatchery. Prog. Fish-Cult. 43:140-141. <br />1025 <br />. 1982. Induced spawning and culture of bon- <br />ytail chub. Ibid. 44:201-203. <br />HOLDEN, P. B., AND C. B. STALNAKER. 1970. $y5[em- <br />aticstudies of the cyprinid genus Gila, in the upper <br />Colorado River basin. Copeia 1970:409-420. <br />Huass, C. L. 1955. Hybridization between fish species <br />in nature. Syst. Zool, 4:1-20. <br />JORDAN, D. S. 1891. Report of exploration in Col- <br />orado and Utah during the summer of 1889, with <br />an account of the fishes found in each of the river <br />basins examined. Bull. U.S. Fish Comm. 9:1-40. <br />AND B. W. EVERMANN. 1896. The fishes'Df <br />North and Middle America. Bull. U.S. Natl. Mus. <br />47(pt.l):1-1240. <br />LAGLER, K. F., J. E. BARDACH, R. R. MILLER AND D. <br />R. M. PASSING. 1977. Ichthyology. 2nd. ed. John <br />Wiley and Sons, New York, New York. <br />MARSH, P. C. 1985. Effect of incubation temperature <br />on survival of embryos of native Colorado River <br />fishes. Southwest. Nat. 30:129-140. <br />MILLER, R. R. 1961. Man and the changing fish fauna <br />of the American Southwest. Mich. Acad. Sci. Arts <br />Lett. 46:365-404. <br />HINCKLEY, W. L. 1973. Fishes of Arizona. Sims <br />Printing Co., Inc., Phoenix, Arizona. <br />1983. Status of the razorback sucker, Xy- <br />r¢uchen texanus (Abbott), in the lower Colorado Riv- <br />er basin. Southwest. Nat. 28:165-187. <br />AND J. E. DEACON. 1968. Southwestern fishes <br />and the enigma of "endangered species." Science <br />159:1424-1432. <br />ONO, R. D., J. D. WILLIAMS AND A. WAGNER. 1983. <br />Vanishing fishes of North America. Stone Wall <br />Press, Inc„ Washington, D.C. <br />SMITH, G. R., R. R. MILLER AND W. D. SABLE. 1979. <br />Species relationships among fishes of the genus Gila <br />in the upper Colorado River drainage. U.S. Natl. <br />Park Serv. Trans. Proc. Series 5:613-623. <br />TYUS, H. M., B. D. BURDICK, R. A, VALDEZ, C. M. <br />HAYNES, T. A. LYTLE AND C. R. BERRY. 1982. Fish- <br />es of the upper Golorado River Basin: distribution, <br />abundance, and status, p. 12-70. In: Fishes of the <br />upper Colorado River system: present and future. <br />W. H. Miller, H. M. Tyus and C. A. Carlson (eds.). <br />Publ. West, Div., Amer. Fish. Soc., Bethesda, Mary- <br />land. <br />VANICEK, C. D., AND R. H. KRAMER. 1969. Life h15- <br />tory of the Colorado squawfish, Ptychocheilus Lucius, <br />and the Colorado chub, Gila robusta, in the Green <br />River in Dinosaur National Monument, 1964--1966. <br />Trans. Amer. Fish. Soc. 98:193-208. <br />VANICEK, C. D., R. H. KRAMER AND D. R. FRANKLIN. <br />1970. Distribution of Green River fishes in Utah <br />and Colorado following closure of Flaming Gorge <br />Dam. Southwest. Nat. 14:297-315. <br />LYNN R. KAEDING, BOB D. BURDICK, PATRICIA <br />A. SCHRADER AND WILLIAM R. NOONAN, U.S. <br />Fish ¢nd Wildlife Service, Colorado River Fishery <br />Project, 529 25%z Road, Grand Junction, Colo- <br />rado 81505. Accepted 21 Nov. 1985. <br />