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~ Page 21 <br />natives cannot survive in the presence of any nonnatives and thus recommend management <br />options to this effect (Clarkson et al. 2005). <br />~ Species Overview <br /> <br /> <br /> <br />i <br /> <br />f <br /> <br />e <br />The three fish species are all in the order Cypriniformes. Roundtail chub (Figure 2-3) are in the <br />family Cyprinidae, the minnows. Flannelmouth sucker (Figure 2-4) and bluehead sucker (Figure <br />2-5) are in the family Catostomidae, the suckers. <br />Common characteristics of endemic, large-river fish <br />including the three fish species, are: 1) fusiform <br />bodies, 2) humped or keeled dorsal surfaces (only <br />present in the endangered Colorado River species, <br />not the three species), 3) leathery skins with <br />embedded scales, and 4) large, often falcate, fins. <br />Such morphological features, combined with <br />relatively long life spans, are thought to be <br />Figure 2-3 Adult roundtail chub (Gila robustal <br />adaptations to the harsh, unpredictable <br />physical environment of the Colorado River <br />Basin (Bezzerides and Bestgen 2002, <br />Minckley 1991, Scoppettone 1988, Stearns <br />1993). All three species reach relatively <br />large sizes (ca. 300 - 500 mm), are <br />relatively long-lived, and are thought to <br />require only sporadic recruitment to <br />Figure 2-4 Adult flannelmouth sucker (Catostomus maintain population stability. Of the three <br />latipinnis) fish species, flannelmouth sucker can <br />demonstrate long-range movement (ca. hundreds of stream miles) throughout the course of their <br />lives, which is not generally observed for the other two species (Bezzerides and Bestgen 2002). <br />The two catostomids are primarily benthic feeders (Banks 1964; Childs et al. 1998; Grabowski <br />and Hiebert 1989; Greger and Deacon 1988; <br />Minckley 1973; Muth and Snyder 1995), whereas <br />adult roundtail chub are frequently omnivorous <br />and can be piscivorous and insectivorous as large <br />juveniles and adults (Bestgen 1985; Karp and Tyus <br />1990; Koster 1957; McDonald and Dotson 1960; <br />Neve 1976; Schreiber and Minckley 1981; ltinne <br />1992; Tyus and Minckley 1988; Vanicek and <br />Kramer 1969). Additional life history <br />characteristics, distribution and abundance have <br />been described in numerous texts and publications <br />for roundtail chub (Bestgen and Propst 1989, Figure 2-5 Adult bluehead sucker <br />Brouder et al. 2000, Voeltz 2002), flannelmouth (Catostomus discobolus) <br />sucker (Chart 1987, Douglas and Marsh 1998, McKinney et al. 1999), and bluehead sucker (e.g., <br />Bestgen 2000, Cavalli 1999, Holden and Minckley 1980, McAda 1977, McAda and Wydoski <br />1983). Bluehead sucker are also discussed in Brunson and Christopherson (2001), Jackson <br />(2001), Mueller et al. (1998), and Valdez (1990). <br />i <br />