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128 <br />MUELLER <br />Minckley et al. 1991). Much effort has gone into <br />research, habitat restoration, and legislative protec- <br />tion. Furthermore, recovery efforts have generally <br />focused on the upper Colorado River basin where <br />the razorback sucker is rare; little has been done to <br />actively manage surviving populations or repress <br />nonnative fish communities. <br />The continual decline of the razorback sucker <br />prompted Lake Mohave field biologists to adopt an <br />active management effort to maintain razorback <br />suckers in Lake Mohave. The group adopted a <br />quick response management approach, rather than <br />developing a more conventional recovery program. <br />The goal was simple: to replace Lake Mohave's old <br />razorback sucker population before the relict pop- <br />ulation was lost. The purpose of this paper is to <br />describe the development, operation, and results of <br />a cooperative program to maintain the razorback <br />sucker in Lake Mohave. <br />Approach <br />Cooperative Partnership <br />The Native Fish Work Group (NFWG) is a co- <br />operative effort of the Arizona Game and Fish De- <br />partment, Arizona State University, Bureau of Rec- <br />lamation, USFWS, National Biological Service, <br />National Park Service, and Nevada Division of <br />Wildlife. The NFWG and the program to save the <br />Lake Mohave razorback sucker were conceived, <br />created, and implemented at the field level. Partic- <br />ipants are local biologists, many directly responsible <br />for managing the Lake Mohave resource. The <br />NFWG has no specific budget; work is accom- <br />plished through the collective resources of partici- <br />pating agencies. The NFWG meets every 4 months, <br />or as needed, to review past work and to plan and <br />assign future activities. Decisions are made at the <br />lowest possible management level. Tasks are as- <br />signed according to available expertise, resources, <br />and function. For instance, environmental compli- <br />ance and permits are usually handled by the Na- <br />tional Park Service, Endangered Species Act (16 <br />U.S.C.A. §§1531 to 1544) compliance is completed <br />by the USFWS, construction activities are managed <br />by the Bureau of Reclamation, and Arizona State <br />University and the National Biological Service assist <br />with research needs. Monitoring activities, fish col- <br />lection, and site maintenance are generally accom- <br />plished by personnel from all the agencies. <br />The NFWG first drafted a research and manage- <br />ment plan in 1990 that outlined the goals and po- <br />tential methods for sustaining the Lake Mohave <br />razorback sucker population. The plan is a working <br />document that identifies and prioritizes goals while <br />allowing for implementation flexibility. The plan is <br />periodically updated as needed. Annual and specific <br />task reports provide information regarding activi- <br />ties. The program has focused resources on actual <br />implementation rather than administrative pro- <br />cesses. <br />Factors influencing the program are limited re- <br />sources, the rapid decline in razorback sucker num- <br />bers, and time. The existing population is dying of <br />old age and could perish by the end of this decade. <br />The program is being expedited by using the repro- <br />ductive potential of thousands of reservoir spawners <br />rather than the conventional culturing practice of <br />mass-producing fish from a small captive brood- <br />stock. <br />Management Concept <br />Habitat degradation in the lower Colorado River <br />basin has been extensive, but research suggests pre- <br />dation is the single most important factor for re- <br />cruitment failure in Lake Mohave (Minckley 1983; <br />Marsh and Langhorst 1988; Minckley et al. 1991). <br />Razorback suckers do successfully spawn in reser- <br />voirs and other lentic bodies of water. Eggs incubate <br />and hatch, but young razorback suckers apparently <br />survive only in environments where nonnative pred- <br />ators are absent or rare (Minckley et al. 1991). <br />Razorback suckers once flourished in several newly <br />impounded reservoirs in the lower Colorado River <br />basin (Minckley et al. 1991). Apparently, popula- <br />tions of razorback suckers were able to expand <br />while reservoirs filled and before nonnative preda- <br />tors became established. <br />The ecology of these fish is not clearly under- <br />stood. However, razorback suckers do spawn with <br />the rising water from spring runoff. Prior to the <br />channelization of the Colorado River and the cre- <br />ation of large reservoirs, larval razorback suckers <br />were dispersed into large, newly flooded, and highly <br />productive nursery areas (Minckley et al. 1991). <br />Survival may have depended on a combination of <br />high spawner fecundity, rapid larval growth, and the <br />dispersal of a naturally small predator community <br />contained by seasonal low river conditions. Today, <br />conditions are much different. Reservoirs have in- <br />undated seasonally flooded nursery areas and have <br />modified, but also expanded and stabilized, aquatic <br />habitats. The early colonization of many of the <br />lower Colorado River reservoirs by razorback suck- <br />ers suggests the fish could tolerate some physical <br />habitat changes. However, it appears the introduc-