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<br />MARSH ET AL.
<br />listing as endangered, efforts began to reintroduce
<br />or augment razorback sucker populations by stock-
<br />ing millions of hatchery-produced and hatchery-
<br />reared larvae and juveniles into the lower Colorado
<br />River and central Arizona rivers from which the
<br />species was extirpated (Minckley 1983; Minckley
<br />et al. 1991; Hendrickson 1993). However, this ap-
<br />proach proved ineffective because predation by
<br />nonnative fishes was extreme and long-term post-
<br />stocking survivorship was nil (Marsh and Brooks
<br />1989; Hendrickson 1993). Largely because of this
<br />program's failure, the razorback sucker was listed
<br />in 1991 as an endangered species (USFWS 1991).
<br />A cooperative partnership was established in
<br />1990 to conserve the razorback sucker in Lake
<br />Mohave, and the group became known as the Na-
<br />tive Fish Work Group (NFWG). Its members in-
<br />clude Arizona Game and Fish Department, Ari-
<br />zona State University, U.S. Bureau of Reclama-
<br />tion, U.S. Fish and Wildlife Service, U.S. Geo-
<br />logical Survey, U.S. National Park Service, and
<br />Nevada Division of Wildlife. The NFWG devel-
<br />oped several research and management strategies
<br />for sustaining razorback suckers in Lake Mohave
<br />but determined that stocking at a size relatively
<br />safe from predation was the only way to ensure
<br />success (i.e., survival) and thereby supplement al-
<br />ternative strategies (i.e., habitat restoration and
<br />nonnative fish removal; Mueller 2003). The
<br />group's goal was to establish a repatriate popu-
<br />lation of 50,000 fish that adequately represented
<br />the known genetic characteristics of the wild Lake
<br />Mohave population that existed when its decline
<br />began. Throughout this paper, "wild" refers to the
<br />naturally produced adults and larvae, and "repa-
<br />triate" refers to fish derived from wild stock,
<br />reared in captivity, and stocked into the reservoir.
<br />Based in part on the work of Marsh and Brooks
<br />(1989), the initial target size at release was 250-
<br />300 mm total length (TL), which required sub-
<br />stantial quantities and reliable sources of larvae
<br />and predator-free rearing environments. One so-
<br />lution was to spawn broodstock maintained at
<br />hatcheries and rear their larvae, but hatcheries
<br />were generally space-limited and cost-prohibitive,
<br />and razorback suckers require up to 2 years to
<br />attain the target size. Hatchery-produced fish also
<br />would under-represent the genetic diversity of the
<br />razorback sucker population in Lake Mohave
<br />(Minckley et al. 1991; Dowling et al. 1996b;
<br />Minckley et al. 2003). The group instead explored
<br />options to produce larvae in seminatural environ-
<br />ments such as lakeside backwaters or offsite
<br />ponds, but early production efforts afforded little
<br />success. That is, approximately 200 wild adult ra-
<br />zorback sucker stocked into a lakeside backwater
<br />before the 1991-1992 spawning season produced
<br />fewer than 300 juveniles, and stripping gametes,
<br />fertilizing eggs, and releasing 200,000 embryos
<br />into a lakeside backwater produced only 17 ju-
<br />veniles from 1991 to 1993 (Minckley et al, 1991;
<br />Mueller 1995). An alternative effort began in 1994
<br />when collection methods were honed to gather nat-
<br />urally produced larvae from inlets and bays around
<br />Lake Mohave (see Mueller et al. 1993). To date
<br />this effort has yielded more than 440,000 larvae.
<br />These larvae were reared in protected lakeside
<br />backwaters and offsite growout facilities that were
<br />free of nonnative predators, and after achieving a
<br />nominal target size, the juveniles were measured,
<br />PIT-tagged (passive integrated transponder; Bio-
<br />mark, Inc., Meridian, Idaho), and repatriated to the
<br />lake.
<br />Monitoring of the NFWG's repatriation program
<br />in Lake Mohave has coincided, to date, with an-
<br />nual assessments of the lake's wild razorback suck-
<br />er population, which began in the early 1970s un-
<br />der the direction of W. L. Minckley at Arizona
<br />State University. This sampling effort has pro-
<br />duced repatriate population estimates that have
<br />been occasionally reported along with the status
<br />of the wild population (e.g., Minckley et al. 2003),
<br />but overall success or failure of the group's goal
<br />to repatriate a population of 50,000 razorback
<br />suckers was not evaluated. However, the program
<br />has achieved success because the genetic charac-
<br />teristics of the wild population have been passed
<br />along from adults to the repatriate larvae (Dowling
<br />et al., 2005). In this paper, we present population
<br />abundance, survival, and growth rate estimates of
<br />the NFWG's razorback sucker repatriation pro-
<br />gram from 1992 through 2002; assess the suc-
<br />cesses and failures of the program from a demo-
<br />graphic perspective; and offer recommendations.
<br />Methods
<br />Once they attain adulthood, repatriated fish were
<br />available for capture as they commingled with wild
<br />razorback suckers on spawning grounds in the
<br />spring. Annual spring (March) censuses of adults
<br />in Lake Mohave coincided with peak razorback
<br />sucker reproductive activity when fish were ag-
<br />gregated in coves and inlets, and along shallow
<br />shorelines, reefs, and spits (Minckley 1983). Less
<br />intensive monitoring also occurred at nonspawn-
<br />ing times of year. Trammel nets (20-100 m long
<br />x 3 m deep, 2.5-3.8-cm inner mesh, and 25.4-
<br />30.5-cm outer mesh) and boat-mounted electro-
<br />fi
<br />tl
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