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BEHAVIOR AND PHYSIOLOGY OF MARKED FISH 653 <br />40so n-n electroshock+marking <br />•- -• single electroshock <br />o• • •O control <br />30 <br /> <br />? 20 - ?u <br />- <br />E <br /> <br />10 bg <br />1 <br />0 <br />0 <br /> <br />E 0 T T <br />0 <br />a 3O <br />U <br />4U 40 00 <br /> 30- T <br /> 20 <br /> <br /> 10 ? Q <br />0 1 3 6 ti <br />T <br />24 168 <br />TIME (h) <br />FIGURE 6.-Mean plasma lactic acid concentrations in unstressed (control) Alsea Hatchery cutthroat trout and <br />in fish subjected to a single 4-s, 300-V-DC electroshock or electroshock plus marking. Upper and lower panels <br />represent first and second replicate trials, respectively. Means (N = 5) within a time interval with no letters in <br />common are significantly different (P < 0.05); time intervals with no letters shown indicate no significant difference <br />among the means. Vertical bars represent 1 SE. <br />might alter catchability (e.g., Saul 1980) and sug- <br />gests that habitat quality is an important variable <br />to consider when conducting population esti- <br />mates. Electrofishing might be inefficient in areas <br />with much instream cover, and another technique <br />(e.g., estimates by snorkeling) may be more effec- <br />tive. Evidence for changes in catchability of <br />marked fish during subsequent electrofishing at- <br />tempts is equivocal (Cross and Stott 1975; Bohlin <br />and Sundstrom 1977; Yundt 1983; Peterson and <br />Cederholm 1984). Given our findings, this effect <br />of electrofishing needs further investigation. <br />Fish seemed to recover faster in sections with a <br />relatively large catch. These sections also had a <br />relatively low percentage of marked fish unac- <br />counted for and were areas where cutthroat trout <br />were found in large groups in deep water. Marked <br />fish often returned to such groups soon after re- <br />lease and were classified as normal fish even though <br />they might still have been affected by the stress <br />and were actually seeking refuge. Section differ- <br />ences in the amount and type of available cover <br />may have affected the observation of marked fish. <br />In areas with deep water and cobble and boulder <br />substrates, it was difficult to maintain position to <br />observe any fish that might have been hiding. Dim <br />light within large boulder complexes and around <br />undercut banks added to the difficulty of observ- <br />ing fish using such areas for shelter. <br />The general lethargy and cover-seeking re- <br />sponse of fish in the artificial stream were consis- <br />tent with observations at Mill Creek, especially