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COPEIA, 1991, NO. 3 <br />130° 120° 110° <br />1 55° <br />1 50° <br />145° <br />1400 <br />135 <br />130 <br />30°1 <br />55°1 <br />50°1 <br />45°1 <br />40°1 <br />35°1 <br />'.0° 110° 1000 <br />° 0 500 <br />km E <br />P. oregonensis <br />a <br />i <br />i <br />- I f r - - <br />' P. umpquae <br />' P. /uc/us i <br /> Fig. 1. Distribution of the four extant species of Ptychocheilus. <br /> ranges from the Nass River, British Columbia, Woahink drainages, and Tahkenitch Lake of <br />h throughout the Columbia River drainage, to western Oregon (Bond, 1980); P. grandis (Ay- <br /> include the Harney Basin of eastern Oregon res) is distributed throughout the Sacramento- <br /> and the Peace River system east of the Conti- San Joaquin drainage of California including <br /> nental Divide in Alberta (Scott and Crossman, the Pit River system, the Clear Lake drainage, <br /> 1973; Wallace, 1980;j. S. Nelson, pers. comm.); and coastal drainages of the Pajaro, Salinas, and <br />+?; L P. umpquae Snyder inhabits the Siuslaw and Russian rivers (Moyle, 1976, 1980a); P. lucius <br />i <br />Umpqua rivers, as well as the Siltcoos and <br />Girard is restricted to the large rivers of the