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MAYDEN ET AL.-SYSTEMATICS OF WESTERN CYPRINIDS 833
<br />eight anal rays for this species, as did Scott and
<br />Crossman (1973) although the last authors were
<br />aware of literature that reported a range of eight
<br />to 10 anal rays. Our data were consistent with
<br />that of Scott and Crossman (1973) who reported
<br />"8, seldom 9" anal rays for P. oregonensis.
<br />1. Anal rays 9, lateral-line scales usually ?80 -__
<br />- -- - -- ------- - - - -- - - lucius
<br />- Anal rays 8, lateral-line scales usually <79 -_ 2
<br />2. Dorsal rays 8
<br />- Dorsal rays 9 - -- - -- -- - - ----- 3
<br />3. Transverse scales 15-18/1/10-13, circum-
<br />peduncular scales 27-30 -_..-_-_.-.--_..---.- oregonensis
<br />4. Transverse scales 19-23/1/12-14, circum-
<br />peduncular scales 32-36 umpquae
<br />Material examined.-Museum acronyms follow Leviton et al. (1985).
<br />HespeniUucus sysmetricus.-Salmon Creek, 2.5 km W of Bodega on
<br />Salmon Creek Rd., Sonoma county, California (UCLA W86-32, N =
<br />15). Mylopharodon conocephalus.-Pit River, 5.5 km W of Canby, Modoc
<br />County, California (UCLA W81-24, N - 15). Plydiecheilm grandis.-
<br />Stillwater Creek, 5 km WNW of Anderson, Shasta County, California
<br />(UCLA W84-21, N = 15); Russian River, Sonoma County, California
<br />(UCLA W86-40, N - 2). Plydweheilus lucius.-Palo Verde, California
<br />(LRCM 745, N - 1); Colorado River S of Mohave Wash, Mohave
<br />Canyon, San Bernardino County, California (CASSU 33212, N = Ir
<br />Delta of Colorado River, United States (?) (CAS-SU 00696, N = 1); at
<br />Yuma, Yuma County, Arizona (CASSU 03833, N = 11 of 15); at
<br />Tempe, Maricopa County, Arizona (CAS-SU 03828, N - 4 of 17);
<br />Green River, Daggett County, Utah (UMMZ 179579, N = 11 of 20
<br />and UMMZ 182540, N - 4 of 6). Ptychocheilus oregonensu.-Umatilla
<br />River W of Echo, Umatilla County, Oregon (UCLA W86-2, N - 15);
<br />Maries River at Kiger Island Rd bridge, Benton County, Oregon (UCLA
<br />W81-17, N = 15 and UCLA W86-3, N - 15); Clearwater River, Mon-
<br />tana (UCLA W70-18, N = 5). Piychocheilw umpquae.-Umpqua basin:
<br />Lookingglass Creek (Umpqua dr.), 0.4 km N of W inston, Douglas Coun-
<br />ty, Oregon (UCLA W864, N - 15); South Umpqua River at mouth
<br />of Myrtle Creek, Douglas County, Oregon (UCLA W81-15, N - 15);
<br />Siuslaw basin: Woahink Creek (Siltcoos dr.), Lane County, Oregon (OS
<br />3333, N = 10); Triangle Lake, Lane County, Oregon (OS 10092, N =
<br />2).
<br />ACKNOWLEDGMENTS
<br />This study was supported by the University
<br />of California at Los Angeles Department of Bi-
<br />ology Fisheries Program. The development of
<br />some of the analytical methods used in this study
<br />was supported by NSF BSR83-07102 and
<br />BSR85-16738 to WJR and DGB. D. L. Swofford
<br />supplied the PAUP program. We have benefit-
<br />ed greatly from discussions with C. Bond and
<br />W. L. Minckley. We thank W. Eschmeyer, J.
<br />Seigel, D. Nelson, and D. Markle for assistance
<br />in examining specimens from the California
<br />Academy of Sciences, the Natural History Mu-
<br />seum of Los Angeles County, the University of
<br />Michigan Museum of Zoology, and Oregon State
<br />University collections. We thank D. A. Carney
<br />and L. M. Page for providing a preprint of their
<br />study and R. R. Miller and J. S. Nelson for their
<br />helpful comments on the manuscript.
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