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<br />Primary successional trends suggest that species with seeds dispersed <br />by wind and water (e.g., willows, cottonwood, alder, and to a lesser <br />extent birch) colonize bare mineral substrates at about the same time. <br />Once these plants are established, environmental conditions such as flooding, <br />drought, grazing or other pressures favor one species over others. When <br />individual plants reach about one meter in height, they attract birds that <br />disperse seeds and berries. This enables many understory shrubs and forbs to <br />become established. Once larger shrubs and trees become established they <br />may persist for some time (for several decades as determined by dendro- <br />chonology of selected tree species) even though the stream may change course. <br />Drier conditions may thin the canopy and favor establishment of understory <br />species that are more subject to fires. Fires alter community composition. <br />It appears that under drier conditions the establishment of cottonwoods, <br />alders, birch, and willow is restricted due to competition from aggressive <br />herbs and grasses and from. lack of adequate soil moisture. <br />Management implications of these findings are that plant establish- <br />ment is a critical factor in riparian plant community patterns. Sites <br />with dry soil surfaces but relatively high water tables (5 to 10 feet) <br />can often support a variety of species such as cottonwood, hawthorn, buffalo- <br />berry, and conifers if these plants can be protected from competition long <br />enough to develop a supporting root system. Other species such as alder, <br />birch and willow are more dependent on the influence of water, either by <br />flood action or inundation, to maintain themselves relatively free from <br />competition. Management strategies to maintain these communities must <br />incorporate artificial or natural manipulation of the stream channel. This <br />is especially true along the drier lower reaches of the stream. <br />If natural reproduction of the mesic species of shrubs and <br />trees is desired, then those managing water diversion projects should <br />recognize the role of the flooding regime and the need for fine texture <br />materials. The use of control or diversion structures that encourage <br />the formation of point bars, or the use of beaver for diversion of the <br />stream channel into smaller flooding channels, will be neceessary to main- <br />tain or enhance riparian vegetation, particularly the alder, willows, <br />and cottonwoods. These species are important in providing stream bank <br />stability, COver for fish and habitat for wildlife. In the past we have <br />mapped these riparian communities but seldom recognized why certain <br />patterns have occurred. The future of riparian systems belongs to those <br />who have the foresight to recognize the dynamic state of flux, the inter- <br />relations of the components of these riparian systems, and who act respon- <br />sibly in their management. <br /> <br />LITERATURE CITED <br /> <br />Hansen, D. J. 1977. Interrelationships of valley vegetation, stream <br />regimen, soils, and solar irradiation along the Rock Creek in the <br />Uinta Mountains of Utah. Ph.D. Dissertation, University of Michigan, <br />179 pp., Ann Arbor, Michigan. <br /> <br />65 <br />