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<br />Table 5. Spawning collections, Colorado squawfish, Green and Yampa rivers. <br /> Number Male Female <br />Year River Total Ripe n TL n TL <br />1983 Yampa 20 19 16 594 3 687 <br />1983 Green 13 12 11 568 1 625 <br />1982 Green 13 7 6 509 1 640 <br />1981 Yampa 34 31 27 535 4 734 <br />1982 Yampa 1 1 1 547 0 <br /> <br />Because so few females were captured, it was concluded that female <br />Colorado squawfish were either difficult to catch or less abundant than <br />males. Since females tend to be larger and older (Vanicek 1969) females <br />may be limited in number compared to males, because of differential <br />mortality. <br /> <br />Spawning Habitat <br /> <br />The locations of Colorado squawfish within the primary (RK 26, RM <br />16.5) and secondary (or replicate) (RK 29, RM 18.2) sites were mapped on <br />each contact day. (Figures 8 and 9). The behavior of these fish was <br />similar to that observed in 1981 (Miller et al. 1982): radiotelemetry <br />indicated the fish were selecting deep pools or eddies, and riffles. As <br />in 1981, the fish would remain in deep pools/eddies, abruptly move to <br />cobble bars, then return to the pools/eddies. This behavior, similar to <br />visual observations made for spawning northern squawfish (Beamsderfer <br />and Bjorn 1982), warranted the division of selected habitats into two <br />apparent types: <br /> <br />1) A resting-staging habitat in pools or large shoreline eddies <br />where the fish may find suitable resting and feeding habitat <br />between spawning forays or where males may gather around females <br />until they were ready to deposit eggs, and; <br /> <br />2) A deposition-fertilization habitat in riffles, where males <br />and females congregate and the females deposit eggs and the <br />males fertilize them. <br /> <br />Our data suggested that spawning Colorado squawfish selected river <br />reaches that had deep pools/eddies in close proximity to riffles. Also, <br />riffles selected for spawning were not uniform, shallow riffle areas <br />that ran all the way across the channel; rather, riffles used lay adjacent <br />to deeper runs. This would seem logical for the Colorado squawfish, <br />since it is a big fish. Should these observations be true, they may aid <br />in explaining why the Colorado squawfish selects the lower Yampa Canyon, <br />and especially the deep pool/riffle/pool habitat between RK 24-30 (RM <br />15-19) for spawning. In the upper canyon, relatively large areas are <br />associated with a different slope-discharge-substrate relationship, with <br />steep slope, high velocity and large substrates (J.S. O'Brien, Personal <br />Communication). <br /> <br />19 <br />