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<br />,.". <br /> <br />Spawning occurred along the river levee shoreline at the same locations where eggs were <br />found last year. Eggs were not visible; however, they were found in the interspaces of <br />small riprap where beaver had been active. The majority of spawning occurred on the <br />central portion of the levee, at the terminus of a large beaver trail and in relatively <br />shallow water (<50 em). The three suspected spawning sites were relatively small in area <br />(<3 m2) and limited to disturbed areas. Selection of previously cleaned substrate is not <br />uncommon for other desert cyprinids (Mueller 1984). <br /> <br />Bonytail appear to be very interactive and social. Smaller fish (10-15 em) were <br />constantly over the spawning area, especially during the night. These fish would form <br />loose schools as they would dart from place to place. Social interactions. generally <br />included darting about, chasing and nudging one another with their snout, and foraging <br />for food. <br /> <br />Bonytai1 densities (video estimates) over known spawning sites averaged 17.44 fish/m2 <br />(3.6 to 55.4, n=222); considering the relatively small spawning area (<9 m2), this <br />suggests that only a small portion (5%) of the community spawned at any given time <br />(2002 estimate: 6,000 bonytai1 > 15 em). Unlike razorback suckers, individual spawning <br />may be shorter in duration (hours versus days) and number of events. It is also quite I <br />possible that we have not discovered all the areas used by spawners. I <br /> <br />It was difficult to determine what was actual spawning because of all the activity. By all <br />indications, spawning was infrequent and rapid. The video recorded two definite <br />spawning acts. The act would often only take 1-2 sec and involved several fish (3-7) and <br />would be followed by the telltale feeding frenzy by several dozen egg scavengers. One <br />event involved a larger, presumed female that was tightly flanked by two smaller males. <br />The three fish dipped in unison to the substrate, with their backs arched. Immediately, <br />two dozen other bonytails aggressively drove into the gravels to feed. The second <br />observation was of a larger group of fish. Again, 4 to 5 smaller males flanked a larger <br />female. They rested their heads in a depression and pressed their bodies together and <br />tails slightly elevated. Unlike the first sighting, these fish finned vigorously for nearly 2 <br />see, which again triggered a feeding frenzy involving about 4 dozen fish. <br /> <br />Other "Factors & Organisms". The videos revealed that razorback sucker were <br />spawning over substrate composed of a mixture of small filles to cobble, while bonytail <br />were spawning over more coarse material, composed of small to large angular rip-rap. <br />This difference in material size may influence survival. Horns and Magnuson (1981) <br />reported that substrate composition was an important factor determining lake trout egg <br />survival. They found that crayfish consumed large numbers of trout eggs. Savino and <br />Miller (1991) showed a correlation between substrate size and survivability of lake trout <br />fry. <br /> <br />We also observed high densities of bullfrog tadpoles and to a lesser extent crayfish <br />feeding among spawning fish. This prompted us to examine their possible role in this <br />pond's ecology. <br /> <br />10 <br />