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<br />15
<br />
<br />taxa between and within the native and nonnative groups (Figure 5). In each year except 1981, native
<br />fishes staned spawning earlier than nonnative species; in 1981, red shiner and fathead minnow started
<br />spawning earlier than all native fishes except Dannelmouth sucker. In each year, all nonnatives except
<br />redside shiner ended spawning later than native fishes. Spawning for both native and nonnative fishes
<br />among years staned and ended earliest in 1981 and staned and ended latest in 1983. Among native fishes
<br />in each year, Oannelmouth sucker staned spawning earliest, foUowed closely by Gila sp., speckled dace, and
<br />bluehead sucker, respectively. Speckled dace ended spawning latest, followed by bluehead sucker, GilD sp.,
<br />and Dannelmouth sucker, respectively. For nonnatives, fathead minnow staned spawning earliest, followed
<br />by red shiner, redside shiner, and sand shiner, respectively in aU years except 1981. Sand shiner
<br />ended spawning latest, followed by red shiner, fathead minnow, and redside shiner, respectively. In 1981,
<br />sand shiner staned spawning earlier than redside shiner.
<br />Correlations among years between values of SPAWN-l and SPAWN-T by taxon were different
<br />between the native and nonnative groups (Figure 5). For all nonnative fishes, values of SPA WN-l were
<br />smallest in 1981 and greatest in 1984 and especially 1983, and values of SPA WN-T were smallest in 1983 or
<br />1984 and, except for redside shiner, greatest in 1981. Coefficients for correlations between these
<br />spawning-period parameters, for all nonnatives except redside shiner, were negative and significant (positive
<br />and moderate for redside shiner). Values of SPAWN-l and SPAWN-T for each native taxon were similar
<br />among years, the greatest difference among years occurring in 1981 with values of SPAWN-I, and
<br />correlations were positive for Gila sp. and flannelmouth sucker and negative for speckled dace and
<br />bluehead sucker; however, correlations were weak for all four fishes.
<br />
<br />Relative abundana.-Values of total (over all reaches) relative abundance from the two indices, i.e.,
<br />catch-per-unit-effort (CPUE) and percentage of occurrence (PO), were positively correlated among years
<br />for each age class of all fishes except fathead minnow (Table 6). Correlations were strong or significant for
<br />age-O speckled dace, bluehead sucker, red shiner, and sand shiner and for both age classes of redside
<br />shiner.
<br />
<br />Variations in total relative abundance of age-O fish among years and relative abundance among
<br />reaches over all years differed mostly between the native and nonnative groups (Tables 7, 8). For all native
<br />fishes except Gila sp., values of total CPUE and PO were greatest in 1984 (greatest in 1981 for Gila sp.),
<br />whereas, for all nonnatives except fathead minnow, these values were greatest in 1981 (CPUE and PO for
<br />fathead minnow were highest in 1982 and 1984, respectively). Smallest values of total CPUE and PO
<br />occurred in 1984 for Gila sp., 1982 for speckled dace, 1981 for blue head and Dannelmouth suckers, 1984
<br />for all nonnatives except fathead minnow, and 1983 for fathead minnow. Among reaches, CPUE for native
<br />fishes was highest over aU years in reaches 1-3 and lowest in reaches 4 or 5. CPUE for red shiner was
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