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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />7 <br /> <br />River near Lily Park (the Uttle Snake River joins the Yampa River at Uly Park; river km 81.8). To place <br />the maximum-daily (peak) discharge of the lower Yampa River for each year during 1980-1984 in historical <br />perspective, values of annual peak discharge in 1934-1991 were taken from Maybell records and frequency <br />and cumulative-frequency distnbutions were prepared. Each peak-discharge value in 1980-1984 was given <br />a percentile rank within the grouped frequency distnbution. <br />Carlson et at (1979) and Haynes and Muth (1982) reported that in 197~1977 and 1981, <br />respectively, estimated peak spawning for most of the more common fIShes in the lower Yampa River <br />occurred sometime during late May through late August Discharge of the lower Yampa River typically <br />begins to increase from over-winter, baseline flows in April, peaks in mid May-early June, remains <br />relatively high through July, and is at or near baseline from August through March. Because river flows <br />exert deterministic effects on various physical conditions important in spawning or survival of young fish <br />(e.g., water temperature, velocity, and quality and formation and maintenance of spawning and nursery <br />habitats), we presumed that discharge conditions during spring-summer would affect annual reproductive <br />timing and success of fishes in the lower Yampa River. Three parameters describing the monthly flow <br />regime of the lower Yampa River during April-July in each year studied were selected for our analyses; <br />i.e., peak (maximum-daily), daily mean, and total discharge. <br />Mainchannel water temperatures of the lower Yampa River typically begin to increase from their <br />over-winter lows in early-mid March, are fairly constant from day-to-day in May-mid June (during <br />spring-runoff discharge), and reach maximum values in early-mid August (after spring-runoff flows have <br />subsided). Degree-days (sensu Arnold 1960; Baskerville and Emin 1969) were used as an index of the <br />annual rate of warming of the lower Yampa River during spring-summer. Degree-days were estimated <br />using mainchannel water-temperature data collected at the Maybell gage in 1981-1984 (1980 data were <br />incomplete). Although this gage is located about 45 km upstream of the study area's upper end, it <br />provided the most continuous data on mainchannel water temperatures (typically, on dates when <br />temperature data were recorded at the Deerlodge Park gage, daily minimum or maximum mainchannel <br />water temperatures were about 1-3oC greater at Deerlodge Park than those recorded near MaybeU). For <br />each year, monthly degree-days were estimated as number of days maximum mainchannel water <br />temperature reached or exceeded 12, 14, 16, 18, or 200C in April-July. These temperature thresholds were <br />selected to represent a range of water temperatures likely affected by variations among years in discharge <br />during spring-summer. It was believed that annual onset and seasonal persistence of each of these <br />temperature thresholds would be especially influenced by magnitude and timing of peak discharges, thus <br />affecting the rate of warming of the lower Yampa River. <br /> <br />Fish daJa.-In the laboratory, samples were first organized by collection date and location (reach)~ <br />Fish from each sample were identified to the lowest possible taxon (most were identified to species; native <br />Gila were left at genus, although most were probably Colorado roundtail chub). Specimens in each taxon <br />