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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />5 <br /> <br />after the water temperature had reached 180C and that late larvae (12-20 mm long) were captured 3-4 <br />weeks later. Results of field studies conducted in the Yampa and Green rivers after 1979 generally <br />confirmed Vank:ek and Kramer's observations and demonstrated that Colorado squawfish spawn in the <br />lower 50 km of the Yampa River, especially in lower Yampa Canyon, from late June through early August <br />(Tyus et a1. 1981; Wick et a1. 1983; Haynes et a1. 1984). Information on principal spawning seasons of <br />other fishes occurring in the lower Yampa River was also considered when planning sampling efforts. <br />Seines (1.0 or 3.0 x 1.2 m; 0.8-mm-square mesh) were used to collect age-O or older fish from <br />backwaters or shoreline habitats with negligtble water velocity (typically s 0.2 m/s; e.g., pools, embayments, <br />and eddies) throughout each reach. Width and length of each seine haul (each seine haul was considered a <br />separate sample) were measured with a metric tape, and area sampled (mZ) was calculated. Samples were <br />collected according to the "qualitative-representative-sample approach" (Hocutt et aL 1974), i.e., all <br />available habitats in each reach that could be seined were sampled to obtain representative collections of <br />fishes that would yield data on relative abundance. Consequently, number of samples co))ected and area <br />seined in each reach per year generally reflected amount of seineable habitat. Water temperature (oC) was <br />measured in each habitat type at time of sampling. <br />Most sampling in 1981-1984 was conducted monthly or bimonthly from early-mid June (shortly <br />after spring flows had peaked) through late September or mid October (Table 2). In 1981 and 1982, <br />samples were also collected in late Apnl (before spring flows had peaked). Collections in 1980 were made <br />in late August and early October. Collections were made primarily during early-mid June through late <br />August in reaches 1 and 2 and in April, late August, mid-late September, or early or late October in <br />reaches 3-5. All samples were fixed in 10% formalin and returned to the Larval Fish <br />Laboratory, Colorado State University, Fort Collins, for processing. <br /> <br />Analyses <br /> <br />Flow antltemperature data.-Various environmental factors influence reproduction of fIShes <br />(DeVlaming 1972; Lam 1983; Bye 1984; Stacey 1984; Munro 1990). Welcomme (1985) reported that for <br />riverine fishes, timing of spawning is related to water flow and temperature and reproductive success or <br />survival of young are affected by the flow regime. Annual discharge and mainchannel water-temperature <br />data for the lower Yampa River were taken from U.S. Geological SUlVey monthly records. Stream gages <br />on the Yampa River were located near the town of Maybell (river km 138.9; about 45 kin upstream of the <br />study area's upper end) and at Deerlodge Park (river kin 72.2; at the upper end of Yampa Canyon). <br />Discharge and temperature data collected at the MaybeU gage were available for each year studied, <br />whereas, collection of these data at the Deerlodge Park gage was started in 1982. Discharge of the Yampa <br />River at Deerlodge Park in 1980 and 1981 was approximated by adding monthly values of daily and total <br />discharge taken from Mayben records to corresponding data collected at a stream gage on the Little Snake <br />