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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />using these parameters/criteria are indicative of viable, self-sustaining populations <br />with successful reproduction, survival and growth. The abundance criteria are <br />intended to surpass minimum standards for genetic viability. Genetic diversity and <br />demographic stability are promoted through the distribution criterion, which <br />requires populations of the listed fish species to be comprised of multiple existing <br />or new population segments. The population abundance criteria are set above a <br />minimum threshold to minimize inbreeding and contribute toward maintaining <br />existing genetic diversity. Recruitment and length frequency distribution criteria <br />are a measure of the dynamic processes of differential annual reproduction and <br />survival of year classes. Recruitment into population segments indicate the <br />movement or dispersal of these year classes throughout their habitat. The <br />abundance trend criterion and the range and frequency attributes of the <br />recruitment and length frequency criteria are important for defining stability while <br />recognizing natural population level dynamics and variability, and for identifying <br />negative trends or conditions for these populations. The use of rolling averages for <br />defining recruitment rate and size category ranges is intended to dampen annual <br />variation in estimation of the criterion due to natural or sampling factors, and are <br />based on empirical data from these same populations demonstrating a persistence <br />and stability over time. Rolling averages are estimated using a proscribed number <br />of sequential annual estimates, and use all estimates within the available time <br />period up to the most recent annual estimate (e. g. 1991-1993, 1992-1994, 1993- <br />1995, etc.). The use of frequencies of a threshold recruitment rate in conjunction <br />with the range of stability is intended to maintain the occurrence of recruitment <br />rates in the higher end of the range of stability. Peaks in recruitment rates are <br />necessary periodically to offset naturally occurring periods of low recruitment, and <br />boost adult population numbers when environmental conditions are favorable for <br />survival through the late-juvenile life stage. This recruitment pattern is considered <br />an adaptive strategy for a long-lived species in a variable environment with <br />uncertain and varying reproductive success. <br /> <br />Distribution of multiple populations and population segments over a wide <br />geographic area provide demographic stability and security from the potential risk <br />of a catastrophic event significantly impacting the species or anyone population by <br />providing redundancy. Maintaining connectivity between these population <br />segments and populations is considered important to allow gene flow between. <br />adjacent populations or permit recolonization. Maintaining connectivity between <br />populations via passage structures or translocation of wild fish around instream <br />barriers is desirable if these barriers cannot be removed. Restoration stocking to <br />accelerate recolonization of vacant habitat is also desirable. In contrast, if <br />augmentation stocking is required to maintain population abundance, this condition <br />is considered a failure to meet a self-sustaining status. <br /> <br />The data available through the Recovery Program centralized database <br />maintained by U.S. Fish and Wildlife Service in Grand Junction have been <br />analyzed for the four endangered fish species. This database is a cumulative <br />collection of all endangered fish data derived from Recovery Program projects <br /> <br />3 <br />