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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />61 <br />can have substantially different feeding histories. Further, length is only an indicator of <br />past growth history, not an indicator of current conditions. Comparing masses of similar- <br />sized fish may provide additional clues to their condition relative to each other. Mass is a <br /> <br />more transient indicator of growth, exhibiting short-term changes depending on <br /> <br />reproductive condition, immediate feeding history, and stress. Proximate analysis of <br /> <br /> <br />body composition, particularly lipids, further provides an early indicator of stress. <br /> <br /> <br />Changes in quantity of reserves such as lipids may be observed prior to any observed <br /> <br />decreases in growth following short-term change in food supply (Rice et al. 1993). <br /> <br /> <br />Energy reserves can furthermore change substantially without affecting overall weight in <br /> <br /> <br />fish. This relationship occurs because water content and lipid content are inversely <br /> <br /> <br />related. A fish gains lipids it loses water and thus may incur no net weight change though <br /> <br /> <br />available energy reserves have increased dramatically (Flath and Diana 1985; Brown and <br /> <br /> <br />Murphy 1991). Lipids may prove particularly useful in analysis of larval fishes because <br /> <br /> <br />reserves are limited and tend to be used rapidly. Due to differences in how each metric <br /> <br /> <br />(length, mass, lipids) respond to feeding history the relationship among them can be used <br /> <br />as a measure of general well-being (Carlander 1977; Wege and Anderson 1978). <br /> <br />The relationship between length and weight is described by the allometric <br /> <br />function: <br /> <br />W = aLb <br />, <br /> <br /> <br />where W is weight, L is length, and a and b are constants. Logarithmic transformation <br />results in the equation: <br /> <br />10gW = loga + blogL , <br />