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<br />25 <br /> <br />Only 18 of 278 recaptured fish had readable scales taken at both capture and r:ecapture. <br />Time interval between capture and recapture ranged from less than I year to 6 years; . Known <br />time interval was compared to observed annuli between capture and recapture. Recapture <br />interval was correctly predicted for seven fish. Ages of these fish ranged from 7 to 13 years at <br />first capture. Intervals of eleven fish were incorrectly determined. Ten of these were incorrect <br />by plus or minus 2 years of the known time interval. One fish, 15 years old, was underestimated <br />by 6 years. A possible source for discrepancies between expected and observed age difference is <br />whether the first true annulus was present on the scale. Another source of variation could be <br />time of year when annulus formation actually took place. It is difficult to identify when annulus <br />formation Occurs on older fish collected between May and July. Incorrectly ageing a scale by <br />only 1 year at both capture and recapture could account for a total difference of up to 2 years <br />from the known time interval. <br /> <br />Age composition was determined for fish from the Colorado, Green, White, and Yampa <br />rivers. Colorado River samples were probably influenced by the small, inadequate sample of 14 <br />fish. The majority of older fish collected from the Green River were age VII, age VIII in the <br />White River, and age IX in the Yampa River (Figure 13). The Yampa River contained a broader <br />range of older fish than any other river. <br /> <br />Scale samples of fish from the Colorado, Green, White, and Yampa rivers were treated <br />both separately and combined for back-calculated lengths (Tables 3-7). Length at first annulus <br />was unknown due to the lack of a first annulus on any scale. The shortest fish length calculated <br />from an annulus was 57 mm. Only 25 fish were calculated to have lengths under 75 mm at the <br />first observable annulus. Some of these may have been the true first annulus, but for most fish, <br />the first annulus was apparently not observed. <br /> <br />Back-calculated fish length was plotted against age for each river (Figure 14). Growth <br />rates of fish younger than 7 years were similar among the four rivers. After age VII or VIII, fish <br />reached sexual maturity based on lengths greater than 428 mm (Table 8) and growth rates <br />differed among rivers. <br /> <br />Musker (1981) used a values from equation (6) that were derived from fish from each <br />river to back-calculate lengths. Due to small samples, these values were not accurate estimates of <br />the body length to scale radius relationship. These estimates of a caused inaccurate calculations <br />of lengths at each annulus. Carlander (1981) warned of the problem of calculating a from <br />inadequate samples and suggested using a standard a for each species. Using a (45.151) calculated <br />from 336 Colorado squawfish (Figure 6), lengths at each annulus were recalculated from annuli <br />measurements reported by Musker (Appendix B). These adjusted calculations were used in <br />further comparisons. <br /> <br />Back-calculated lengths (this study) were shorter than lengths at age reported by other <br />studies (Table 8). Fish tended to be 1 year older for a given size when ~ompared with lengths at <br />age from previous studies (Figure 15). The oldest fish aged by Musker (1981) was an <br />879-mm-TL, 13-year-old Yampa River fish. An ll-year-old female, 610 mm long was the <br />oldest fish aged by Vanicek and Kramer (1969). Seethaler (1978) also aged his oldest fish at lJ <br />years. The two oldest fish I aged were from the Yampa River and were 18 years old and over <br />800 mm long. Oldest fish from other rivers were 16 years (Green and White rivers) and 12 years <br />(Colorado River). <br /> <br />A verage annual growth increments were determined from back-calculation table <br />summaries (Tables 3-7; Figure 16). Growth increment between years was greatest (over 60 mm) <br />between 3-4 or 4-5 years. Yearly growth decreased to about 30 mm per year after age XII, but <br />tended to increase in later years (approximately Age XVI). Relative growth calculated from <br />back-calculation tables decreased with age to between 5-10%, and fish over 12 years grew only <br />