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<br />0' <br /> <br />.' <br /> <br />of the potamon zone has decreased as a consequence of the downstream extension (discontinuity) <br />of the rhithron zone through regulation of flow from the deep storage reservoirs. The concept of <br />ecosystem "resets" and discontinuities (sensu Ward and Stanford 1983), coupled with the notion <br />that connected channel and floodplain (backwaters, wetlands) components of the riverscape are <br />seasonally pulsed by flooding (Ward 1989), robustly integrate the myriad of biophysical processes <br />that are influenced by stream regulation. Strong inferences about how a river ecosystem may <br />respond to alternative flow management actions must be derived in this ecosystem context. The <br />downstream shift in the position of the rhithron-potanlon transition is an ecosystem-level measure <br />of change wrought by regulation and should be used as to adjust flows to maximize conditions <br />known to be favorable to both potamon (e.g., endangered fishes) and rhithron (e.g., trout) fisheries. <br />7) Strong food web interactions are likely occurring as a consequence of the presence of a <br />wide variety of nonnative fishes, which now dominate fish communities throughout the Upper <br />Colorado River Basin. Despite obviously important effects of predation and competition for food <br />resources, very little written information exists about the ecology of nonnatives in the Upper <br />Colorado River Basin. High volume flows appear to reduce numbers of nonnative species, and <br />diversion dams installed many years ago (e.g., Redlands on the Gunnison River) may have <br />segregated nonnative populations and limited range expansion. However, much more information <br />is needed. I suspect that considerable unpublished data exists in files as a consequence of the <br />sampling effort required to collect significant numbers of the endangered fishes. If so, the <br />information should be examined relative what is known about the native species, and it should be <br />published. If not, sampling protocols should be developed to describe trends in nonnative <br />populations in all of the river segments. In addition, experiments (e.g., as described below in <br />Section VI) are needed to clarify interactions between natives and nonnatives. <br />8) River ecosystems are too complex to be described by simple detenninistic models or <br />constructs of individual attributes. Ecosystem components are N-dimensional, inherently variable <br />(stochastic) in time and space and interact in complex ways that cannot be predicted from simple, <br />logistic equations. Construction of an ecosystem model that describes all of the dynamic processes <br /> <br />44" <br />